Thursday, April 7, 2016

Arthur D Hasler Professional PhotoBio

! Buy the new book !
The Way Home:
A Photo Biography of AD Hasler
by A Frederick Hasler
now available on line…... from the link:

from the createspace store for $23.70

Tributes to Arthur Davis Hasler and The Way Home, A Photo
Biography by his son Fritz (Arthur Frederick Hasler)

Arthur D. Hasler was one of the foremost limnologists of the 20th century, but there are many more elements to his rich and fascinating life: beyond his work in limnology and animal behavior and his University of Wisconsin career, Hasler was a Germanophile who served America in Germany in World War II, a family man, and a lifelong Mormon.  His son Arthur F. Hasler’s new book The Way Home uncovers many aspects of his multidimensional life and recounts them with a spectacular collection of large, high quality black & white and color photographs.

Lynn K. Nyhart,
Vilas-Bablitch-Kelch Distinguished Achievement Professor
Department of the History of Science, University of Wisconsin-Madison

In my book The Dancing Bees - 2016, the story of Arthur D Hasler is woven inextricably into the life of Karl von Frisch. It is wonderful to see their interaction so richly illustrated through photographs and archival documents in The Way Home, the 2017 book by his son Arthur F. Hasler.  Hasler’s contributions to science were significant, and he continues to hold a special place with family, friends, students, and colleagues who remember a life well-lived.

Tania Munz, author of The Dancing Bees, Karl von Frisch and the discovery of the Honeybee Language
VP for Research and Scholarship, Linda Hall Library of Science, Engineering & Technology, Kansas City, Missouri.

As Arthur Davis Hasler’s last graduate student, I would like to describe what it was like to work with this scientific giant of a man and convey what his 53 Ph.D. students felt about him. Herr Doktor Professor Hasler, or more affectionately “Doc” as he was called by the cadre of students with whom I was associated, was unanimously appreciated for both his scientific rigor and helpful advice that he freely gave to us. We admired and respected him. He prepared us so well, and all his students were so highly thought of by the scientific community, that within 3 months of our graduation, almost all of us had either a teaching or research position at another university. 

Fritz’s book captures the entire life of Arthur Davis Hasler and provides rare sagacity into how and why scientific discoveries are made. The book also presents Arthur Davis Hasler family man and community/political activist. Finally, it provides insight into the central role of the University of Wisconsin in developing the science of limnology.

I thought Fritz did a wonderful job of integrating his dad’s scientific accomplishments with his family history. I was particularly impressed with Chapter 7 that covered Doc’s role in the U.S. War Department’s 1945 Strategic Bombing Survey in Germany and Austria. Excerpts of his letters to his wife and children were illuminating, capturing the extent of the devastation wrecked upon the allied bombing of these Axis countries. The chapter also describes Doc’s meeting with Nobel Laureate Karl von Frisch, who discovered the dance language of honey bees, whereby bees communicate the direction and distance to sources of nectar to other bees in the hive. In a letter home, Doc included a description of witnessing one of von Frisch’s experiments, which for me was the highlight of the book.

Allan T. Scholz,
Emeritus Professor of Biology (Fisheries), Eastern Washington University
Cheney, Washington    May 5, 2017

 A remarkable photographic portrayal (and tribute) to a remarkable scientist, Arthur D. Hasler. This unique, thoughtful and loving memoir fills in much important information about one of the key leaders in the long and important history of limnology at the University of Wisconsin-Madison. A must read for anyone interested in the history of aquatic ecology in general and especially of limnology at the University of Wisconsin. Many aspects of Art’s life and scientific achievements are reported in this book. It will serve as an excellent reference document for Limnology in Wisconsin during the Hasler era.

Dr. Gene E. Likens, 
University of Connecticut, Special Advisor to the President on Environmental Affairs
Cary Institute of Ecosystem Studies, Founding Director and President Emeritus, Millbrook, New York.
Led a team that discovered Acid Rain, 1963
Elected a member of the National Academy of Sciences, 1981

National Medal of Science Award winner, presented by President George W. Bush, 2001

From the UW Center for Limnology
What an amazing and wonderful tribute to Arthur Davis Hasler. The Way Home, a photo biography created by his oldest son, Arthur Frederic (Fritz) Hasler captures both the profession and the family of Arthur, the scientist and the father. The Way Home hints at the importance of his family and details Art’s major contribution of learning how migrating salmon find their way home to spawn. Arthur, like many scientists and fathers, had these two families -- his genetic family of parents, children, and their descendants, and a professional family of scientific programs, graduate students and colleagues, and their descendants. Arthur was devoted to both families. He succeeded admirably in both roles. Today Arthur Davis Hasler is appreciated and honored by each of these important populations.
Clearly, the author and son, Fritz, loved Art, and he honored, respected, and admired Art as a professional scientist and educator. These elements permeate the photo selection and brief text that accompanies each photo. Our perception is that Fritz learned a great deal about his scientist father as he discovered photographs and new aspects of Art’s life. For Fritz this has been a process of collecting his and the family’s memories and discovering new revelations about Art that took place after he, Fritz, had left home. Many members of Art’s genetic family helped with photos and memories. Similarly, members of Art’s professional family helped to find photographs and contributed their memories. Archives maintained by the University of Wisconsin were important to this endeavor. Fritz’s enthusiasm and skill with visual images had developed as a professional analyst of satellite images with NASA for 30 years.

This, then, is a juxtaposition of Art’s personal family life with his professional life as a scientist and educator. It becomes obvious that both are related and intertwined through time. Professional colleagues will discover the family behind the man and what appeared to have driven his approach to science and its application to human concerns about water organisms and ecosystems. The family will discover, as did Fritz, the contributions that Art made to doing cutting-edge science, educating the next generations of scientist and educators, building and fostering scientific institutions, and addressing local to global environmental and scientific challenges.
John J. Magnuson, Founding Director of the Center for Limnology (Emeritus)
James F. Kitchell, Director of the Center for Limnology (Emeritus)
and Stephen R. Carpenter, Director of the Center for Limnology
University of Wisconsin-Madison

May 2, 2017
Professor Arthur Davis Hasler, PhD
Zoologist, Limnologist, Ecologist

University of Wisconsin Madison

Professional Photo Biography by son Fritz Hasler

This Blog Post is sponsored by a member of the Hasler Family

A series of photos and documents of Art's professional life, arranged mostly chronologically with explanations and commentary by me, other family members and colleagues. 

Sources: Art's family color slide collection, Limnology Laboratory Archive images, Power Points by John Magnuson, his first wife, Hanna's, family photo album, University of Wisconsin Archives, The negatives of Art's photographs from Europe in 1945, Art's lecture slide carousel, digital photos that I took after 2000, related images from the internet, and a biography of ADH by Gene Likens (included at the end). Various family photos. often taken by Art, are added for context. John Magnuson (Emeritus Director of the Limnology Lab, David Null (Director of the University of Wisconsin Archives) Daughter Sylvia and son Galen have been very helpful in collecting and scanning photographs. All of Art's children have been helpful in assembling material, adding anecdotes, and making corrections.

Art, Lago Maggiore, 1955
(Arthur Hasler Photo)

Biographical Synopsis

From my memories, Art's genealogy, and a biography by his student, Gene Likens

Note: the Gene Likens biography and a biography by his last student, Allan T. Scholz appear in a separate post in this blog

Arthur Davis Hasler was born January 5, 1908, in Lehi Utah to Walter Thalman Hasler and Ada Elizabeth Broomhead. Arthur married Hanna Bertha Prusse on September 6, 1932, in Provo Utah. They were married for 37 years until Hanna died in 1969. Art and Hanna had six children: Sylvia, born 1936, Arthur Frederick (Fritz) 1940, Bruce Davis, 1942, twins, Galen Rolf and Mark Rudolf 1945 and Karl Gregory, 1947. Arthur married Hatheway Minton, July 24, 1971 in Madison Wisconsin and was married to her for 30 years until his death in 2001.

Arthur learned to play the French horn in high school and continued to play throughout his life including 20 years for the University of Wisconsin Symphony and the Madison Civic symphony. Arthur was very active in the Boy Scouts of America and received the Eagle badge in 1924. Arthur served a 30 month mission for the LDS (Mormon) Church to Germany, Czechoslovakia and Austria, from July 1927 to February 1930. He learned the German language very well on his mission and continued to study and love it for the rest of his life. He graduated from BYU in 1932 and married Hanna September 6 1932 in SLC. They left directly for the University of Wisconsin in Madison when Art began his graduate studies in Zoology. In 1935 he took a job with the US Fish and Wild Life Service on the Chesapeake Bay and worked for two years while living in York Town Virginia, where Sylvia was born. 

He returned to the University of Wisconsin in 1937 to finish his PhD in Zoology and begin as an instructor in the department. He was promoted to Assistant Professor in 1941, Associate Professor in 1945, and Full Professor in 1948. He served on the Strategic Bombing Survey in Germany for four months in 1945 for the U.S. Army Airforce. 52 doctoral students (a full deck) and 43 masters students received their degrees under his supervision  His research study that brought him the most acclaim was the scientific proof that salmon learn the odor of their birth stream as smolts (fingerlings) and find their way back to the that same stream 1.5 to 5 years later as adults by the use of the sense of smell. He worked on and refined that study from 1945 through the end of his career in 1983. He wrote and coauthored over 200 peer reviewed scientific papers and authored or contributed to seven books. In 1954 and 1955 he went on a one year sabbatical as a Fulbright Research Scholar at the University of Munich Germany with famous honeybee researcher and Nobel Prize Winner, Professor Karl von Frisch.

He wrote the proposals and received funding from the National Science Foundation to build the Laboratory for Limnology at the University of Wisconsin in 1963.  He was elected as a member of the National Academy of Sciences in 1969. In American scientific circles this is an honor short only of winning the Nobel prize. He also received numerous other awards and honorary degrees from other Universities. He received funding for another research lab, this one on his beloved Trout Lake in Northern Wisconsin. He retired in 1978. In 2006 he received the post humus honor of having his lab renamed the Arthur D Hasler Laboratory of Limnology. He survived four different kinds of cancer including, colon, neck, and lung starting in 1972. The three malignant cancers of the colon, lung and lymph glands were treated by undergoing two major surgeries and chemotherapy. He died March 23, 2001 of old age.

Hundreds attended his funeral in Madison.

Figure 1: Walter T and Ada Hasler, Provo 1933

(Hanna & Art Family Album)

Arthur Davis Hasler was born on January 5, 1908 to Walter Thalmann Hasler and Ada Elizabeth Broomhead in Lehi Utah.

WT Hasler was the son Swiss immigrants. His father was a musician who brought music education to the small town of Mt Pleasant in north central Utah.

Walter was the first in his family to pursue higher education. He attended the Baltimore College of Physicians and Surgeons (Now the University of Maryland School of Medicine) received his MD degree, and practiced as an Eye, Ear, Nose, and Throat Specialist in Provo Utah for many years.

Figure 2:  Hasler Children, Provo, 1915

(Hanna & Art Family Album)

Bill, Ada, Art (7), Thalmann, Calvert just after having moved from Lehi, to Provo Utah.

Art's oldest brother Calvert died of diabetes at the age of 16 in spite of his doctor father's best efforts in the days before insulin was discovered.

Art's older brother, Thalmann (Tommy) went medical school and became a doctor like his father.

Art graduated from college at the height of the depression and with his father suffering from cancer, there was not enough money in the family to send him to medical school like his brother. 

As a result, Art applied for a graduate assistantship at the University of Wisconsin in the Zoology Department, and the rest is history (see Figure 9 etc.)

Figure 3: Art with other Eagle Scouts, Provo, 1924
(Hanna & Art Family Album)

Middle row right wearing Eagle Pin on his left pocket at age 16 (see the pin on his eagle sash - Figure 183)

Figure 4: Art, Provo, 1926
(Hanna & Art Family Album)

Art age 18, a High School Graduate

Figure 5: Art, Provo, 1927
(Hanna & Art Family Album)

Art age 19, Freshman at BYU Provo


Art's LDS Mission to Germany and Austria
 where he learned German

Figure 6: Art, On his 30 month LDS Mission, Germany, 1928
(Hanna & Art Family Album)

Art served a two-and-one-half year mission for the LDS (Mormon) Church in Germany, Czechoslovakia, and Austria from 1927-29. 

Dad worked diligently studying German and learned it well. He loved German and the German people and defended them even when Germany wasn't popular in the U.S.

I don't know how many times I heard him say this:

"Meine Beziehung zu der deutschen Sprache ist zu meiner Frau, ich kenne sie, ich liebe sie, aber ich beherrsche sie nicht."

I dictated it to my iPhone using the German keyboard that makes-up for my poor German spelling.

English translation: "My relationship with the German language is the same as the one I have with my wife. I know her, I love her, but I don't rule her"

Art returned to BYU in the Fall of 1929 and graduated with a BS in Zoology in 1932.  His study of Utah high Uinta mountain lakes during his zoology studies got him interested in limnology.

Figure 7: Arthur Hasler, Passport, 1927
(UW-Madison Archives)

His son Galen found this in the University of Wisconsin Archives along with a complete record of his correspondence through the years. From the stamps on the passport we know that Art landed in Liverpool England on July 23 1927, served his mission in Germany, Czechoslovakia and Austria. This passport expired May 23, 1929, but it must have been extended and he probably returned in February of 1930 for a total time served of 30 months.  Also note that at 21 he gave his height at 5' 10" He must have matured very late to 6'. In his letters from Salzburg and Linz Austria in 1945 he recalls serving there 15 years earlier on his mission.

Figure 8: District President Hasler, Vienna, Jan 1929


"Pres Hasler des Öestereichischen Distrikts"
(President Hasler of the Austrian District)

Professionally dad was somewhat embarrassed by this part of his resumé. When asked about his mission, he would emphasize the work he had done in Germany to promote the Boy Scout program of the LDS Church. However as you can read in this history, you can see that dad served in very important leadership positions on his mission and could have emphasized that part of his service.

He was the Branch President (minister) for a small congregation near Dresden in East Germany.

He was also the District President (chief ecclesiastical administrator) for all the Mormons in Vienna Austria and for Breslau East Germany (now part of Poland) during different periods on his mission. 

Figure 9: President Hasler, Colleagues, Breslau, October 1929 

By this time Art was District President for the 2nd time, now for the city of Breslau and surrounding   See him here (First row 2nd from left)with the President of the German Austrian Mission, Hyrum W Valentine, visiting from Dresden. 


Art comes back to Utah and finishes his BS Degree in Zoology


Art Marries Hanna Prusse and goes to UW Madison for Graduate School

Figure 10: Art marries Hanna Prusse, Provo, 1932.
(Hanna & Art Family Album)

Formal Wedding picture: Hanna Bertha Prusse and Arthur Davis Hasler

If you knew Art professionally, you also knew how important his wife Hanna was to his professional career.

Art and Hanna were married on September 6, 1932 in the historic Salt Lake Temple. Her sisters Irmgard (left center) and Eveline (right), who were tiny girls on the immigrant boat (from Hanover Germany to Galveston) with her in 1913, were her maid of honor and one of the bridesmaids. Art's brother Bill was the best man.

Figure 11: Art's Graduate School Application, Madison, 1932
(UW-Madison Archives)

Hanna and Arthur were married September 6, 1932, in the Salt Lake Temple and they left immediately for graduate school in Madison, Wisconsin”

Figure 12: Newlyweds Art and Hanna, Milwaukee, 1933. 
(Hanna & Art Family Album)

  This is the first picture we have of the newlyweds in Wisconsin. It was taken in front of the LDS Church in Milwaukee, in 1933 when they were both 25.

Figure 13: Art & Hanna, Madison ,1933
(Hanna & Art Family Album)

Art and Hanna arrive on the University of Wisconsin Campus from Utah
 (Bascom Hall in background)

Art and Hanna were married for 37 years from 1932 until Hanna's death in 1969 at the age of 61.

Figure 14: Hanna, Old Lake Lab, Madison, 1933
(Hanna & Art Family Album)

Hanna on the pier ready to go fishing with Art on Lake Mendota.

Figure 15: Art's Fishing License, Door County, 1933
(Fritz Hasler family heirloom)

Even the future fish professor had to have a fishing license to fish legally in Wisconsin.  By 2016 I think the price of a fishing license, even for In-State Residents had increased a bit.

Figure 16: The Big Three - Birge, Juday, Hasler

(Cover: Transactions of the Wisconsin Academy of Sciences)

Birge, Juday, and Hasler pioneered Limnology at the University of Wisconsin Madison.

Cover of Transactions of the Wisconsin Academy of Sciences, Arts & Letters - Special Issue Breaking New Waters A Century of Limnology at the University of Wisconsin - 1987.  The book recounts the history of the work by Birge, Juday and Hasler.

Edward A. Birge  1851-1950 (Life span 99 years)
Chancy Juday 1871-1944 (Life span 73 years)

Arthur D. Hasler 1908-2001 (Life span 93 years)

Figure 17: Birge, Juday, Trout Lake, 1930
(Hanna & Art Family Album)

Edward A. Birge and Chancey Juday at the Trout Lake Research Station. 

Birge found time to work with his students after retiring from his responsibilities as President of the University of Wisconsin Madison (1918-1925). Birge, Juday, and Hasler were members of the Zoology department at the University that is housed in Birge Hall.

However, in Art's oral history, he describes Birge at this point in his career as flitting in for a couple of days, a few times during the summer, driven in by a UW limousine, probably a perk he retained as a former president of the University. 

Figure 18: Birge, Crystal Lake, 1930
(Hanna & Art Family Album)

Birge near Trout Lake, with the "sun machine" on top of his car

The sun machine was probably a reference device for his measurements of light penetration into the water (see below)

Figure 19: Birge, Baum, Trout Lake, 1933

(Hanna & Art Family Album)

Birge and Hugo Baum, with the "sun machine" just down the shore from the Trout Lake Camp

According to Art's oral history, Birge is measuring light penetration into the water.

Figure 20: Art, Baum, Trout Lake, 1933

(Hanna & Art Family Album)

Hugo Baum and Art building "Lime Floats" at the Trout Lake Camp.

Arthur D. Hasler, the third member of the Big Three is just a graduate student working with Juday at this point. He won't receive his PhD until 1937 and won't become a full professor until 1948.

I'm guessing that Baum is sitting on a finished "Lime Float" designed to release lime from the barrel slowly into the water.

Figure 21: Trout Lake Camp, 1933

(Hanna & Art Family Album)

Figure 22: Trout Lake Camp, 1933

(Hanna & Art Family Album)

Students,Trailer, Boats, Motors

Today in  2016, the pier and the cabins of the Trout Lake Camp on South Trout have been gone for decades, but the beach is now a public picnic area that anyone can visit. Subtract the pier, boats and trailer and it looks 83 years ago very much like it does today.

Go North from Woodruff on 51, turn right on M, turn left at the first Trout Lake Forestry Head Quarters entrance. Just beyond the bike trail on the left is a place you can turn in and park. Continue 200 yards on the path down to Trout Lake.

Figure 23: Art, Juday, other Students, Trout Lake,  1933
(Hanna & Art Family Album)

Summer of 1933 at the Trout Lake Research Station in Northern Wisconsin

Art top, second from left, Chauncy Juday bottom, third from left

Front row from left: Hugo Baum, Edward Schneberger, C. Juday, Sam X. Cross, Militzer and William Spoor

Back row from left: Ray Lanford, A. D. Hasler, Robert Hunt, V. W. Meloche, and Harold Schemer.

Identifications by Mrs. Juday 1956

Figure 24: Juday, Art, Birge, Students, Trout Lake, 1934

(Hanna & Art Family Album)

Professors Edward A. Birge, and Chauncy Juday, with graduate students including Art Hasler third from the right. Chauncey Juday is in the center in the dark coat. Art studied Lake Mendota from the shores of the UW campus and the lakes in Northern Wisconsin as part of his graduate studies.

From the left: David Frey, Martin Baum, John Schreiber, Don Kerst, Harold Shomer, C. Juday, E. B. Fred, Richard Juday, A. D. Hasler, Paul Pavcek and E. A. Birge.

Figure 25: Putting in a boat, Trout Lake, 1934
(Hanna & Art Family Album)

Trailing a boat was a little more primitive in those days, but they already had the idea. Note: Outboard on the car windowsill, and the oars on the running board. 

Figure 26: Fyke Nets, Trout Lake Camp, 1934
(Hanna & Art Family Album)

Fyke Nets were used for catching yellow perch,


Art leaves school and takes a job with the US Fish and Wild Life Service in York Town VA

Figure 27: Art, Oyster Apparatus, Yorktown, 1935
(Hanna & Art Family Album)

Art with a an instrument he devised for measuring the amount of water an oyster processes through it's body.

Art had left the University of Wisconsin in 1935 before he finished his dissertation, to take a job for two years with the US Fish and Wild Life Service in York Town, Virginia 

Figure 28: Art, US BF Lab, Washington DC, 1936

(Hanna & Art Family Album)

Art doing Glycogen Analyses at the U. S. Bureau of Fisheries Lab in Washington DC during his two year stint in York Town VA.

Figure 29: Prusses + Arthur & Sylvia, Provo,1937. 
(Hasler Family Album, scanned from Bill Prusse Negative)

Spring 1937: The first grandchild joins the Prüsse Clan. 

Art, Hanna, & baby Sylvia upper right.

Art's wife Hanna was the eldest of 13 children. Her father, baker Wilhelm Prusse, brought his wife and the first five children from Hanover Germany to SLC in 1913 just before the start of WWI

In this picture, the Haslers had just made the long trek west to Utah from Yorktown Virginia to attend a Prusse family reunion. They probably came by car, taking over a week on the road with a stop in Madison, in the days before freeways. They brought their first child, Sylvia, born October 4th 1936, and the first grandchild of Wilhelm & Johanne to meet them and Sylvia’s 13 aunts and uncles.

Figure 30: Art, Rangers, Crater Lake NP, 1937
(Hanna & Art Family Album)

Art and other Rangers at Crater Lake National Park in Oregon.

It's all making sense now. Art keeps heading west to take a summer job on the west coast as a ranger and Hanna stays behind with her folks in Utah with one-year-old Sylvia.

Figure 31: Ranger Art, Big Fish, Crater Lake, 1937
(Center for Limnology Archives)


In the Fall he started his job as an Instructor at the University of Wisconsin in Madison and finishes his PhD.

Figure 32: Chauncy Juday, Madison, 1937
(UW-Madison Archives)

Art gets his PhD  in 1937 on the physiology of digestion of plankton Crustacea (including Daphnia) from Chauncy Juday

Figure 33:  Art, Madison Civic Symphony, 1938
(Hanna & Art Family Album)

Art, back row right with the French Horn to the right of the right bass player.

When he is not working for the US Government in Virginia, the National Park Service in Oregon, working on his PhD, teaching students, doing his research, raising a family, or driving his family to Utah, Art begins his 20 year career playing the French horn with the Madison Civic Symphony and the University of Wisconsin Symphony.

Figure 34 Art, with French Horn, Madison
(Hanna & Art Family Album)

Art's whole family was musical, practically the von Trapp family singers.
This picture was taken in 1953, inserted here to show Art close-up with his French Horn.

Back Row: Galen, Bruce, Karl, Hanna, Fritz, Art
At the Piano: Sylvia, Mark

Hanna helped dad in so many ways throughout his career. That included performing for his professional guests

Figure 35: Art, Birge Hall, Madison, 1940
(Hanna & Art Family Album)

Probably Birge Hall.


Art Joins the US Army Strategic Bombing Survey and Spends 5 months in Germany and Austria

He Meets Nobel Prize Winner. Karl von Frisch, in Austria

Figure 36: Art with Colleagues & Jeep in Alps, June1945
(Arthur Hasler Photo)

Figure 37 Art & Colleague Somewhere in Europe May1945
(Arthur Hasler Photo)

In 1945, just before the Nazis surrendered, Art volunteered for the Strategic Bombing Survey with the US Army in Germany. Using his German language skills he interviewed German civilians to determine the effectiveness of the indiscriminate bombing and complete destruction of German cities including civilian populations by high explosives and firebombing during the war.

Today we would call this "a crime against humanity".

The Survey determined that the the bombing of civilian populations did not break their will, and may even have strengthened their bond with the Nazi government.

The US went on firebombing Japanese cities, including Tokyo.

The Nazis had started bombing civilians in Guernica Spain(subject of Picasso's famous painting) during the Spanish civil war and continued on with aircraft and missile bombing of civilians in London during the battle of Britain. Therefore it is not surprising that the allies responded with bombing of civilian populations in Germany. Astonishingly, no one repeated the horrible chemical warfare from WWI.

After a search of family archives (where we found his letters from Germany), the Limnology Lab, the UW-Madison Archives at the Steenbock Library, we finally found Art's negatives from his photographs of Europe in 1945 in an off-site Limnology storage facility in Stoughton. We have still not found his photo album from the trip.

On May 7, 1945 Art landed in Paris on VE day as a volunteer..... (Documented in his letter to Hanna. "The whole plane wondered why Paris was crazy celebrating". They could not make London due to weather and arrived the next day when London was celebrating. "The English waited a day" he commented in his letter) The timing is another amazing happenstance of his lucky life.

Figure 38: Darmstadt Following RAF Firebombing, 1944
(Sept 1944, Found on the Internet)

Art arrived in Europe on VE day, May 8, 1945. We were finally able to locate the photos he took and we have the almost daily letters he wrote home from May through August 1945. He spent his first night in Paris and then witnessed the VE celebrations on May 9th in London the next day. He spent the first 6 days in London getting trained and outfitted then traveled two days by plane and jeep to his first duty station in Darmstadt Germany.

Darmstadt had been firebombed during RAF raids on the night of the 11th and 12th of September 1944 that decimated the city and killed over 14,000 residents. So when dad arrived there eight months later, he got to see the results and hear firsthand the stories of those terrible days.

Figure 39: Corpus Christi Procession, Munich, 11 June 1945
(Found on the Internet)

Frauen Kirche and battered Munich.

Art went through Munich only 10 days earlier and must have seen the central city much as it appears here. Miraculously the towers of the iconic Munich Frauen Kirche survived the bombing. 

He saw firsthand the terrible destruction of Darmstadt and Stuttgart and must have seen what we see above. We know that von Frisch expected that his house in the suburbs would be safe even if his lab at the Universität München built by the Rockefeller Foundation was in danger. Art visited the partially destroyed lab on June 2nd. von Frisch moved his extensive library on animal physiology to his home. Unfortunately his house suffered a direct hit, The house and library were completely destroyed except for a milk can hanging in a nearby tree. von Frisch moved to his family home in in the tiny town of Brunnwinkl Austria just east of Salzburg and continued his work.

Munich  was substantially rebuilt as a result of the Marshall Plan and the Wirtschaft's Wunder (economic miracle) by the time our family spent 12 months there 10 years later in 1954 and 1955. I recall the main nave of the Frauen Kirche and numerous buildings in Munich were still in ruins or under construction. Building cranes had sprouted all over Munich. We did see the famous Trümmerfrauen (rubble women) in action in a few locations pulling brick after brick out of the ruins and stacking them for reuse.

30May1945, Starnberg (just south of Munich),  Dear Hanna & Kiddies

Art was in Darmstadt for about two weeks before being transferred to Innsbruck and Salzburg Austria via Munich

Art traveled by Jeep from Darmstadt to Stuttgart and on to Munich. He was amazed by the German Autobahns, the first modern superhighways (the term Art our present Interstates) that did not exist in the US at the time. Much like in the TV Series Band of Brothers, he traveled south on the pavement in his small jeep convoy while legions of POWs and displaced persons, of every nationality were straggling up the median strip and road shoulders in the opposite direction pushing baby carriages, pulling carts, riding heavily loaded bicycles etc. Everything would go smoothly until they hit a big Autobahn bridge or overpass that the Nazi's had blown up to impede the allies, At that point a temporary winding gravel bypass would lead traffic down into the valley across the river on a pontoon bridge and back up to the Autobahn.  Many of these detours still existed 10 years later in 1954-55 when our family spent a year in Germany.

On the outskirts of Munich he saw the sign to Dauchau and was fully aware of Nazi treatment of political prisoners and extermination of Jews there. (I visited Dauchau again with my Grandson in 2014). Art reports that the Germans he interviewed were aware of Dachau as a prison camp for political prisoners, but were not aware of the Jewish extermination there and in Auschwitz (in Poland) and the other foreign camps.

Art describes his visit to Hitler's Eagle's nest on June 11th only 35 days after it had been liberated by the US 101st Airborne (Band of Brothers) - See photo in a separate blog post.

Figure 40: Darmstadt in Ruins, May 1945
(Arthur Hasler Photo)

17May1945, Darmstadt: Dear Hanna and Kiddies,
        (Mostly in Art's handwritten words)

He was billeted in the villa of a former Nazi "bigwig" who had poisoned himself, his grown daughters and their children before the US troops arrived. He slept on a canvas cot under two army blankets and was fed by transient German women. He was still being trained as an interviewer.

"On the way to work we pass along a street where not a single house stands complete" Even so there are Germans living in the city....I don't know where, cellars no doubt. Everyone's clothes are pressed and clean inspite of the dust and rubble surrounding them. 

They look well, but no one is fat. Their ration is one wiener-sized piece of meat a week. Professor Kirkpatrick is still with me along with Riegel (German Language Professor from UWand Workman.

In London, I learned to drive a Jeep and have a license. Also got dog tags like a regular GI

Had a women to interview today with a 3 year old blond boy like Bruce (Art's son of the same age). Sure made me homesick for my little boys (Fritz 5, Bruce 3, Galen & Mark babies), Sylvia (age 9, and my Hanna.

It is too depressing, the tales of these people whether formerly rich or poor... tears at my heart strings. Kiddies are so hungry for sweets and need them."

22May1945,  Darmstadt, My dear Hanna (now typewritten)

"In this city there are not only deaths from the front but deaths from bombing. This large city was burned and blasted completely in a 35 minute raid last September. Thousands were burned in their cellars. He relates the story of a man who watched his daughter running towards him, but was consumed by the fire before she could reach him. Inspite of this terror, it is surprising that it did not break their resistance and more surprising that they were able to go about their work within a few days.

It will soon be a month since I left home--- still no mail. I have written every other day since I left. It makes the time go so slowly when I don't hear from you.

I am still convinced that you need to differentiate between the 80,000,000 Germans and the 2-4,000,000 Nazi Party Members."

"This large city was burned and blasted completely in a 55-minute raid last September. Thousands were burned in their cellars. In spite of this terror it is surprising that it did not break their resistance and more surprising that they were able to go about their work within a few days "  

Figure 41: Kids in Rubble, Germany, 1945
(Arthur Hasler Photo) 

This picture Art took, best captures the spirit of his stories of arriving in Germany just after WWII had ended.  The three little girls are standing with one of Art's colleagues in front of a huge pile of rubble. Each girls is holding something round, an apple maybe? Art always had chocolate and gum for the kids, but not usually fruit. One girls is holding a milk can. When we lived in Munich for a year in 1954-55 we had cans just like that one. There was no prepackaged milk in bottles or cartons, You would take the can to the Milcherei (milk shop), the shop keeper would place it under the spigot and pull down a big lever until the can was full.

Art arrived in Germany just after many German cities had been totally destroyed by allied bombing. Art shared his K-rations and chocolate with the children. He walked down a street with virtually all buildings destroyed on either side (see Darmstadt description above). Then out of a basement under the rubble, a little girl emerged, perfectly dressed ready to go to church. I didn't see this exact story in his letters, but remember it from "sitting on his knee" at age 5.


Figure 42: Art, Salzburg, 1945

(Salzburg Photo Studio Photo)

Major Hasler, US Army Uniform, taken in a professional photo studio in Salzburg Austria

Art was stationed in Salzburg from June 16 to June 27 1945. Once again extreme luck strikes Art. He is stationed only 27 km and 30 min from Brunnwinkl where his hero, Professor Karl von Frisch, is studying his honeybees.

Figure 43: Art,von Frisch, Wolfgangsee, Austria, June 1945
(Arthur Hasler Photo)

Karl von Frisch and his colleague are wearing the traditional Tyrollean Lederhosen (leather pants (shorts)) worn today mostly by children and waiters in restaurants.

Karl von Frisch, Art Hasler, Brunwinkl, 1945 

Galen found this in dad's old lantern slide collection in the Center for Limnology's deep storage facility in Stoughton WI.

His letter of June 17 1945, Art describes traveling from Salzburg only 27 km and 30 min to St Gilgen, Wolfgangsee and Brunnwinkl to visit his idol Professor Karl von Frisch. In a three page typewritten letter he describes in detail, meeting von Frisch and observing him working with his honeybees. 

Figure 44: von Frisch & Frau,Wolfgangsee, Brunnwinkl, 1955
(Arthur Hasler Photo)

17June1945 Salzburg, Dear Hanna and Kiddies

The picture above of von Frisch was taken 10 years later in 1955, but in this letter Art describes traveling from Salzburg only 27 km and 30 min to St Gilgen, Wolfgangsee and Brunnwinkl to visit his hero, Professor Karl von Frisch. In a three page typewritten letter he describes in detail, meeting von Frisch and observing him working with his honeybees. He had only a beehive, a stop watch, a pan of sugar water and 5 colors to dob on the thorax of the bees. With these primitive tools, his imagination and keen sense of scientific observation he learned enough about bees to win the Nobel Prize.

Figure 45: Letterhead, Nazi Stationary, Innsbruck, June 1945
(Arthur Hasler letter to Hanna)

Looking closer, I see this is a Nazi sticker on UW Madison stationary. Other letters from Innsbruck were on embossed Nazi stationary.

Art was mainly stationed in Innsbruck and Salzburg. Some of his letters from Innsbruck are written on Nazi Stationary. By August 8th he was in London on his way home, he is in NYC on the 10th and after a few days with his sister in NY he was on a plane to Madison. On August 18th he wrote a letter from Madison to his 9-year-old daughter Sylvia who is still in Utah describing his final days in Europe.

Figure 46: Art, Sylvia, Fritz, Madison, 25August1945
(Arthur Hasler Photo)

Art in his army uniform. He had just returned from the war, on August 15.

Sylvia had just returned from two months in Utah with Art's folks and I (AFH) had just celebrated my 5th birthday on August 21


Back at the University of Wisconsin in Madison
Art starts work on Olfactory Imprinting and Homing of Salmon

Figure 47: Art, Students Madison, 1950
(UW - Madison Archives)

Art and watching his students taking perch out of a gill net on Lake Mendota

Doing fish population studies.

Figure 48: Art, Photometer, Madison, 1950
(UW - Madison Archives)

Art with a photometer on Lake Mendota.

Figure: 49: Lab Boat, ONR Impulse, Madison, 1952

(Arthur Hasler Photo)

Grad students socializing on Lake Mendota

Figure 50: Art, Schmitz, Ragotzkie, Madison, 1953
(Arthur Hasler Photo)

Graduation day - Schmitz gets his Masters and Ragotzkie gets his PhD, University of Wisconsin.

Schmitz would get his PhD from Art in 1958

Figure 51: Homing Salmon, Hasler, Larson, August 1955
(Scientific American Cover)

The Homing Salmon, Hasler and Larson, August 1955

Seminal article in the popular literature on the homing of salmon. Larson, is not a limnology colleague, but a professional science writer that Art hired to make sure that the article, intended for the general public was readable.

Figure 52: Minnow training tank
(Center for Limnology Archives)

A diagram from the article showing how the aquaria were used to ask minnows the question: can you smell very dilute quantities of chemicals and differentiate between water from different lakes?

Figure 53: Art, Wisby, "Weird Science", Madison, 1947
(Center for Limnology Archives)

One of numerous aquaria (Figure 49) that Art used in the old Lake Lab. He used them  to prove the sensitivity of fish to the smell of minute quantities of chemicals in the water. 

See old lab at the end of Park Street in Figure 12

Figure 54: Art, Wisby, "Weird Science2", Madison, 1947
(Allan T Scholz, Volume 1 Fishes of Eastern Washington)

Figure 55: Art, Warren Wisby, Madison, 1950
(UW - Madison Archives)

Warren observing the minnow responses and Art recording the data. 

Figure 56: Art, Sarles, Editors, Madison, 1950
(UW - Madison Archives)

Art with the rows of aquaria used to access a fish's ability to smell, in the old Lake Lab on Lake Mendota at the end of Park Street on the UW campus.

Two Wisconsin daily newspaper editors are shown as they tour the UW Lake Laboratory. Will Conrad, Medford Star News editor, and Charles E. Broughton, Sheboygan Press editor, listen to Professors William B. Sarles and Arthur B. Hasler of the UW Zoology Department talking about fish and lake water.

One of two images. Published in Wisconsin State Journal November 4, 1950.

Figure 57: Diagram of Homing Research Tank
(Homing Salmon, Hasler, Larson, August 1955)

Art's ingenious tank for proving that salmon "home" using the sense of smell. Notice the miniature fish ladders going up the four arms. The next step was to prove it in an actual river system. (Taken from Center for Limnology Archives)

Figure 58: Homing Tank, Madison, 1952
(Center for Limnology Archives)

This is a photograph of the homing tank shown in the diagram in the previous figure.

Off to the State of Washington for an experiment in nature with some big logistical hurdles.

Figure 59: Wisby, Homing Tank, Madison 1952
(Center for Limnology Archives)

Wisby looking into the central holding compartment of the homing tank.  Odors are released into the arms. The minnows are trained to swim up the ladders in one of the arms to be fed in response to the odor that is released. The dilution of the odor is continually increased to determined the sensitivity of the minnow's sense of smell.

Figure 60 : Seattle and Issaquah Creek Wiers, 1952
(Imprinting &  Homing of Salmon...1983 Hasler/Scholz book)

Map from Wisby/Hasler 1954 Article showing the location of the weirs on Issaquah Creek and the East Fork of Issaquah Creek, East of Seattle Washington. The map shown here is the simplified version used in the 1983 Hasler/Scholz book.

Recalling Art's genius to capture salmon above a fork in a river, take them below the fork, stuff the noses of half of them and release them, capture them again above the fork to prove that only the salmon without stuffed noses could tell which way led to the stream of their birth. 

You might guess that Wisby was very instrumental in the whole experiment since he is the first author on the paper. I don't think dad farmed it out to Wisby because he was too lazy to write it up.

This experiment is described in the article:

W.J. Wisby, & A.D. Hasler. (1954) Effect of olfactory occlusion on migrating silver salmon (O. kisutch). Journal of the Fisheries Research Board of Canada, 472-478

If any of the rest of you lay people were wondering what O. Kisutch means:

Species: Okisutch. Binomial name. Oncorhynchus kisutch (Walbaum, 1792). The coho salmon (Oncorhynchus kisutch; Karuk: achvuun) 

Another Interesting Note: A previous experimenter with the same idea in1926, and had severed the olfactory nerves of the altered fish.

However, apparently the fish were so impaired by the surgery that they wouldn't do much of anything.

61: Issaquah Creek Weir Locations
(Found on the Internet - Text added by AFH)

Figure 62:  Area 17 miles East of Seattle
(Google Maps Text added by AFH)

 Location where the experiment was conducted

Figure 63: Weir Locations, Issaquah Creek
(Google Maps Text added by AFH)

Weir 1 at the Issaquah State Salmon Hatchery on Issaquah Creek is still in use today although the town of Issaquah has probably built up substantially since 1953 

Interestingly the location of Weir 2 - on the East Fork of Issaquah Creek is now in the median of I-90

Figure 64: Wisby, East Fork Issaquah Weir 2, 1952
(Arthur Hasler Photo)

This is the weir 2 (fish trap) built by Art and Warren Wisby on the East Fork of the Issaquah Creek to catch salmon. The salmon were then carried down stream 1.5 miles below the confluence and released. Half the salmon had their noses plugged with cotton. Salmon on the main Issaquah Creek were captured at the Issaquah State Salmon Hatchery (see Figures 64-68).

Figure 65: Wisby, Art, Hanna & Kids, Issaquah, 1951
(Arthur Hasler Photo)

Issaquah and East Fork Confluence

Warren Wisby and Arthur Hasler looking at the confluence of the East Fork of the Issaquah and Issaquah Creek. Art's daughter Sylvia remembers this trip. The women seated is likely Art's wife Hanna, with son Bruce seated beside and Fritz wading in the creek.

I believe downstream is to the left. So Warren and Art would have released the captured salmon in the main Issaquah Creek to our left.  The salmon would swim upstream from our left. Of the 27 with unplugged noses that were captured on the smaller East Fork, 19 would repeat the left turn into the smaller East fork while 8 would "mistakenly" continue up the main Creek. Of the 51 salmon with plugged noses only 12 would make the "correct" left turn into the East Fork while 39 would "mistakenly" continue up the main creek

Figure 66: Tagged Salmon, Issaquah Creek, 1952
(Arthur Hasler Photo)

Control Salmon, tagged, but with unplugged nose, ready to be released below the confluence

Figure 67: Wisby, Issaquah Creek, 1952
(Arthur Hasler Photo)

Warren releasing red salmon below the confluence

Figure 68: Results from Wisby Hasler '54 Experiment.
( University Course Viewgraphs:

Top half: Results from the experiment.

You can see that the unaltered salmon captured on the main Issaquah Creek, on release continued 100% up the creek while none made the left turn up the East Fork.

Of the unaltered salmon captured up the smaller East Fork, on release 70% made the turn up the East fork while 30% continued up the main creek.

Of the salmon captured in the main Issaquah and then had their noses stuffed, on release 76% continued up the creek while 24% made the "false" turn up the East Fork.

Of the salmon captured in the smaller East Fork and then had their noses stuffed, on release 84% "falsely" continued up the creek while only 16% made the "correct" turn up the East Fork.

Although the sample is small, the control group (unaltered) had a very high success rate making the same choice they had made originally, while the altered group (stuffed noses) had a much lower success rate.

Can you imagine the logistics of building the new weir across the whole East Fork, capturing 302 salmon, transporting them down stream, tagging them, stuffing half their noses and releasing them without injury. Then they still had to capture them again to get the results of the experiment.

Bottom Right: A picture of Konrad Lorenz who managed to imprint goslings as hatchlings on himself instead of the mother goose, Result - they are following him around rather than their mother.

Figure 69: Stuffing Salmon Noses, Issaquah, 1953
(Arthur Hasler Photo)

Issaquah Creek, Washington State, from the 1954 Wisby/Hasler article

Original B&W image was found in Art's classic lecture slide carousel


As brilliant and elegant as the salmon nose stuffing experiment was, it had several major deficiencies:

1) The sample was small and would need to be repeated to be conclusive.
2) It was not conclusively known whether the salmon were using long term memory from imprinting years earlier as smolts or only short term memory from the part of the stream where they were just captured.
3) It was not conclusively known whether the salmon had been born in these streams or were perhaps straying.
4) It was possible that the nose stuffing had incapacitated the salmon in other ways besides preventing them from using their olfactory capabilities.

For this reason Wisby in a paper he wrote in 1951 proposed an experiment that would involve imprinting baby salmon as smolts with an artificial chemical and attempt to lure them to a stream independent of their birth location. No one took up the challenge until the late 1960s when salmon were introduced into the Great Lakes and Hasler and Scholz set out to conduct the experiment.  This experiment is reported later in this blog post.

Figure 70: Salmon, Nose stuffed with cotton, Madison.
(Fritz Hasler Photo)

The only color picture I have at present, of a Salmon with a stuffed nose.

From the Limnology Lab Naming Ceremony in 2006 (see Figure 196) 

Figure 71: Issaquah State Fish Hatchery, Issaquah, 1951
(Arthur Hasler Photo)

Figure 72: Wier, Issaquah State Fish Hatchery, 1951
(Arthur Hasler lecture slide carousel)

The hatchery in Issaquah Washington on Issaquah Creek served as Wier 1 for the Hasler/Wisby experiment.

The photos below show the hatchery more recently.  

Figure 73:  Salmon Jumping, Issaquah Creek Hatchery
(Found on the Internet)

Salmon Jumping, Issaquah Creek Hatchery , Washington Sate

Salmon jumping up small dam at the Issaquah State Salmon Hatchery, 17 miles east of Seattle on Issaquah Creek. Salmon swim up the Issaquah creek and are caught in the Hatchery. In the Hatchery, they are killed for their eggs and sperm, which are used to hatch more salmon.

Figure 74: Issaquah State Salmon Hatcher, Issaquah Creek
(Found on the Internet)

Note: Sluice gate/fish ladder on right where some salmon climb right into the hatchery to be caught (see next photo)

Figure 75: Catching Salmon, Issaquah Creek Hatchery,  
(Found on the Internet)

Half the salmon in Wisby and Art's experiment described in the 1954 article were caught at this Washington State Salmon Hatchery on Issaquah Creek.

Figure 76: Salmon below dam, Issaquah Creek Hatchery
(Found on the Internet)

Note: One salmon on far left trying to jump the dam - Issaquah State Salmon Hatchery on Issaquah Creek

Figure 77: Salmon Spawning
(Found on the Internet)

Salmon that make it back to their birth stream turn red as a result of exposure to fresh water.

Figure 78: Salmon Spawning
(Found on the Internet)


Back at the University of Wisconsin in Madison

Figure 79: Art, Fisherman, Madison, 1954
(UW - Madison Archives)

Limnology research wasn't just summer fun! Art and his crew also did a lot of research in the winter.

Art with a local fisherman during a creel (population) census on Lake Mendota about 1954

Fishing for perch through a hole in the ice is a popular winter pastime in Wisconsin.


                      Art's Public Service Work

Figure 80: Proposed Parking, Wisc S Journal, Madison, 1954

(UW - Madison, Archives)

Article in the Wisconsin State Journal on a proposal to construct a 470 car parking lot on fill on the shore of Lake Mendota just west of the Memorial Union. Art vehemently opposed the plan which would have been located right where the new Limnology Lab was eventually built in 1962.

In addition to his work killing the proposed parking lot in Lake Mendota, Art work tirelessly over many years to cleanup the Madison lakes.

Madison is located on an Isthmus between Lakes Mendota (see above) and Monona.

When Art arrived on the UW campus as an instructor in 1937, Lake Monona and Waubesa (the lake below it in the chain) were a cesspool, totally infested with algae due to the fact that Madison sewage had been pumped directly into lake Monona. Mendota was better, but was often overgrown with weeds due runoff from farmers fields and the effluent from two smaller upstream towns. 

Art worked throughout his career, to stop the poisoning of the algae with copper sulphate, and getting the sewage from Madison and upstream towns routed around the lakes.

Figure 81: Algae Bloom, Yahara River, Madison, 1945
(UW - Madison Archives)

Before Art and others got the sewage rerouted,  the lakes down stream from Madison; Monona and Waubesa, and the Yahara River connecting them, were a cesspool, totally infested with algae due to the fact that Madison sewage had been pumped directly into lake Monona until 1936 and Waubesa until 1958.

SourcePerspectives on the eutrophication of the Yahara lakes by Richard C. Lathrop  (Published online 29 Jan 2009) at

Figure 82: Young lady entangled in weeds, Madison, 1954
(Arthur Hasler Photo)

Art was a master of public relations. Here he uses an attractive young women to dramatize the disgusting nature of the weeds growing in eutrophicated (too many nutrients) lakes.

Figure 83: Weed harvester, Madison
(Center for Limnology Archives)

One of the weed harvesting machines that Art advocated using rather than poison to deal with the nutrient rich lakes around Madison.

Figure 84: Weed Harvesting on Madison Lakes
(Found on Internet)

Figure 85: Hasler Boys Whaler, Lake Mendota, Madison
(Arthur Hasler Photo)

Art's photo of the family boat showing that Lake Mendota still had a problem with algae blooms in 1963.

The waste water from upstream communities was not diverted until 1971. Even after the diversion, eutrophication is still occurring due to runoff from fertilizer from lawns and farmers fields.

SourcePerspectives on the eutrophication of the Yahara lakes by Richard C. Lathrop


Art takes Fulbright Professorship Year in Munich Germany and takes the whole family along

Figure 86: Art with Professor von Frisch, Munich, 1954.
(Center for Limnology Archives)

Art takes a Fulbright professorship in Munich and brings the whole family of eight to Germany for a year.

Karl von Frisch was the Nobel Prize winning honey bee researcher that Art had met in Austria in 1945. From family memories: von Frisch's children said they would have starved to death if Art hadn't smuggled food to them right after the war. By now, Art had figured out that salmon find their way back to their birth stream, using their sense of smell. At this point he was trying to find out how the salmon find their way in a huge ocean, back to the mouth of the correct major river. Navigation using sun or stars?

Figure 87: von Frisch, Brunnwinkl, 1954
(Arthur Hasler Photo)

von Frisch abandoned his home in Munich during the War and moved back to his family home in Brunnwinkl Austria on beautiful Wolfgangsea to avoid the heavy bombing of the city. Art found him here when he looked him up when he came as part of the Strategic Bombing Survey in 1945. Art kept up the relationship sending food at first, then helping von Frisch obtain chemicals and scientific equipment that was not available in Europe right after the war. von Frisch is wearing the hearing aid that Art got for him when he found out that he was hard of hearing.

Figure 88: von Frisch and Lorenz, 1955
(Magnuson View Graph)

Konrad Lorenz was another of dad's colleagues that he visited during the year in Munich. The Nobel Prize in Physiology or Medicine 1973 was awarded jointly to Karl von Frisch, Konrad Lorenz and Nikolaas Tinbergen.

Figure 89; Konrad Lorenz, Geese Imprinting, 1955
(Arthur Hasler Lecture Slide Carousel Photo)

Figure 90: Haslers, Munich, August 1955.
(Arthur Hasler Photo)

Karl (named after Karl von Frisch), Mark, Galen, Bruce, Fritz, Sylvia, Hanna, Art

Along with his professional career, Art was a husband and the father of 6 children. Here is the whole family as they were leaving Munich after Art had been doing a year of research with Professor von Frisch.

Haslers “Departure from Munich” in the back yard of the Kunigundenstrasse 55, house in Munich Germany just before they returned to the States in August of 1955. Note: family members are wearing large pretzels around their necks and three of the boys are wearing Lederhosen (leather pants)


Back at the University of Wisconsin in Madison

Fig. 91: Art, Wisby, Colleague, Homing Tank, Madison, 1957
(UW-Madison Archive)

Figure 92: Art, Braemer, Homing Tank, Madison, 1957
(UW-Madison Archives)

In this picture, Art is talking to visiting scientist Wolfgang Braemer about the homing tank shown in this picture at the lab on Lake Mendota in 1957. I remember dad having a similar tank in Munich in 1954, but it was indoors and he used lamps to simulate the sun.

Figure 93: Old Lake Lab, Madison, 1957
(UW-Madison Archives)

This is the lab that Art used with his graduate students when he came back from Virginia in 1937 until the construction of the new Limnology Laboratory in 1962. We see it here, at it's peak, with a big platform on the end of the pier for the big fish homing tank (see Figure 84 above) and other tanks. 

The memorial Union is on the left with Hoofers student recreation club canoes on the shore. The temporary quonset hut on the right is one of numerous huts built all over the UW campus during WWII in the 1940s. The last ones were not replaced with a permanent buildings until the 1990s. The zoology department used quonset huts for many years.

Figure 94: Art, with Students, Madison, 1957
(UW-Madison Archives)

Art, Ragotzkie, Horall, von Frisch, Wisby, Madison, 1957

Bob Ragotzkie with SCUBA gear on ladder, Ross Horall with earphones, Otto von Frisch (son of Karl) with flipper & SCUBA gear, Warren Wisby in Cabin, Art holding light cord.

Art got significant grants of money and equipment from the Office of Naval Research (ONR) the boat, the IMPULSE was a surplus Naval vessel supplied by ONR.  Sonar gear, and probably the SCUBA gear came from ONR as well.

Figure 95: ONR Impulse, Old Lake Lab, Madison, 1957
(UW-Madison Archives)


Art and Colleagues begin Whole Lake Experiments

Figure 96: Art with Bill Schmitz, Saw Mill Pond, 1956
(Center for Limnology Archives)

Trying to use air bubbles to turn over a lake on the Guido Rahr Property on the Wisconsin/Michigan border. Now part of the Tenderfoot Forest Preserve.

Figure 97: Art with colleague Schmitz, Saw Mill Pond, 1956
(Center for Limnology Archives)

Figure 98: Schmitz, Saw Mill Pond, 1956
(Arthur Hasler Photo)

Figure 99: Hanna, Boys, Schneider, Madison, 1956
(Arthur Hasler Photo)

Hanna, Hasler boys, VIP Zoology dinner guest, Hans Schneider from Germany, eating Christmas turkey, Hasler home 205 Lathrop St.

Schneider wrote a paper on Grunion sounds related to migration.

Hanna served dinner to a countless number of Art's visiting guests at the Hasler home in Madison.

Art is the photographer taking the picture with his Leica camera and flash.

Figure 100: Liming Lakes, Northern Wisconsin 1933, 1956
(Magnuson Viewgraph)

 Liming Lakes over the years, Northern Wisconsin 1933, 1956

On the left, Art with Hugo Baum building lime floats at Trout Lake in 1933

Figure 101 Art's son Fritz,  Liming Peter Lake, 1956
(Arthur Hasler Photo)

Classic Limnology Lab picture on how to change the chemistry of a lake. Identical Paul lake was left untouched as a control.

Figure 102: Fritz, Liming Peter Lake, 1956
(Arthur Hasler Photo)

That's me above  (Art's eldest son Fritz) emptying a 50 lb bag of lime into a basin, I'm about 16 so this is actually about 1956. In Figure 92 on the left, Art with Hugo Baum building lime floats at Trout Lake in 1933. I'm thinking that both methods have the same goal, but obviously doing it with a high volume motorized pump will get dissolved lime into the water a whole lot quicker.

Figure 103: Fritz, Liming Peter Lake, 1956
(Arthur Hasler Photo)

In this picture you can see a tub outside the boat where Fritz is dumping the lime. A big white hose carries the limey water to the pump that shoots it out the back of the boat

The driver is operating a 5 hp outboard motor to move the boat.

Figure 104: Peter & Paul Lakes.
(Found on the Internet)

Peter & Paul Lakes that "looked like an hour glass or a pair of spectacles" with a narrow passageway between them.  Art got permission to use a bulldozer to make one lake into two. Experiments were conducted on Peter (right?) and Paul (left?) remained untouched as a control.

Although Art worked tirelessly to protect the Madison Lakes from effluent nutrient pollution, copper sulphate poisoning of algae, and land fill, he was not afraid to "mess with mother nature" to advance scientific knowledge of lake systems. 

The liming of lakes shown above, turning over lakes with compressed air shown earlier, the introduction of radioactive radioisotopes in the 1960s,  the acidification of lakes with sulphuric acid by his successors in the 1980s were all examples of the Hasler whole lake experiment approach.

 He used bulldozers to divide one lake into two and to make ponds in the UW Arboretum for experiments. At one point he even dynamited the UW Arboretum in a failed attempt to make a pond.

You might say he used nature like a kid using a chemistry-set for doing experiments. Most of these experiments could never be approved under current EPA and DNR rules and will never be repeated. 

In his defense, these were all small scale modifications of nature and will have no lasting significant effect.

Figure 105: Three of Art's sons, Trout Lake Camp, 1958
(Arthur Hasler Photo)

Karl, Galen, Mark

This was one of the historical Trout Lake Camp cabins used already in the 1930s but now long gone since the building of the new Trout Lake Lab in the 60s.

Figure 106: Art, Lab Coat, Madison, 1958
(Center for Limnology Archives)

Almost luminescent!

Figure 107: Art, Tibbits, Large Mouth Bass, Madison, 1958
(UW-Madison Archives)

Big fish in a small tank

Figure 108: Art, Students, Old Lake Lab, Madison, 1960
(UW-Madison Archives)

Figure 109: Art with Hawaii Dolphin 1960
(Center for Limnology Archives)

Figure 110: Art, Passport Photo and Stamps, 1961
(Center for Limnology Archives)

Art, the well traveled Scientist

Figure 111: Anderegg, Art, Stewart's Dark Lake, 1969
(UW-Madison Archives)

John Anderegg and Art preparing "hot" radioisotope for insertion into the lake.

Figure 112: Art, Anderegg. Likens, Stewart's Dark Lake, 1960
(Breaking New Waters/Center for Limnology Archives)

John Anderegg, Gene Likens, and Art holding a scintillation counter used for the detection of isotopes in a study of the movement of radioactive nuclides from the bottom of a stratified lake. 

Paraphrasing from a conversation with Gene in 2016: "We were investigating the implications of a proposal to put radioactive waste in deep ocean trenches. This lake was very stratified and didn't turn over. We put these dangerous hot isotopes on the bottom of the lake. Coming back later we found organisms on the shoreline that had ingested the isotopes on the bottom, brought them to the top and were still very "hot". It would have been a very bad idea to put nuclear waste deep in the ocean........

At one point when I was taking the hot radio isotopes out of their lead container and was about to put them into the water, Art said "hold a second while I take a picture" however, not wanting to overexpose my self to radiation, I put them immediately into the water."

Gene Likens wrote the biography of Art that is included at the end of this blog post. 

Likens became a professor at Dartmouth College and Cornel University and was later the director of the New York Botanical Gardens. He is famous for discovering Acid Rain and received the National Medal of Science from President George W. Bush

Likens was born in 1935


Art Hosts the 15th International Conference of Limnology at the University of Wisconsin in Madison

Figure 113: Art, Ragotzkie, Madison, 1962
(UW-Madison Archives)

Lake Mendota, Picnic Point in background

Prior to the 15th International Congress on Limnology, Profs. Robert A. Ragotzkie (left), local chairman, and Arthur D. Hasler (right), executive chair of the congress, inspect yellow lotuses on Mendota Bay near the willows. 

Figure 114: Art, Hosting the SIL Meeting, Madison, 1962
(Center for Limnology Archives)

Art, Hosting the International Society of Limnology Meeting, Madison, 1962

This was the 15th SIL and the first time it was held in the US.

UW Student Union Terrace, with Hoofers boats in background.

Paraphrasing from Art's oral history: "I got a grant from the National Science Foundation to sponsor this event. It allowed me to hire an Executive Director for a year and to charter a flight from Europe to bring over the best scientists. European scientists had to apply for the travel grant and we picked the ones we thought would make the biggest contribution to the meeting."

Figure 115: Art, Ragotzkie, Float Plane, Madison, 1962
(Center for Limnology Archives)

Art was always using the latest technology, You could use a float plane to get into a lake in Northern Wisconsin that you couldn't access otherwise. Probably Mendota on Picnic Point, considering the UWZ boat on the left.

Figure 116: Art, Ragotzkie, Float Plane, Madison, 1962
(UW-Madison Archives)

The float plane belonged to the Department of Meteorology. 

Art frequently worked together with departments beyond his home Zoology Department.  

His son Fritz received his PhD from the UW Meteorology Department in 1971 and went on to a 30 career at NASA Goddard Space Flight Center. This blog post is a retirement project of Fritz, where he has put his image processing and computer skills to use.


Art Builds the Laboratory for Limnology at UW Madison and Trout Lake Station

Figure 117: Laboratory for Limnology, Madison, 1962
(Found on the Internet)

(Now Arthur D Hasler) Laboratory for Limnology

Art wrote the proposals and received funding from the National Science Foundation to build the Laboratory for Limnology at the University of Wisconsin in 1963.

Once the bids came in for the unique cantilever design, he was very disappointed that the size of the lab had to be reduced to meet budgetary constraints.

Figure 118 Hasler Limnology Lab, Madison, Spring 1962
(Arthur Hasler Lecture Slide Carousel)

This picture of the lab from behind with the lake in the background was one of Art's favorites that he kept in his standard lecture slide carousel.

Figure 119: Boat Basin, Limnology Lab, Madison
(Fritz Hasler Photo)

A great place to keep the boats indoors in the new lab. (taken 2006 at the naming ceremony)

The Hasler boys were the first to buy a Boston Whaler. They teamed up to buy a 12' 2" Boston Whaler with a 50 hp Mercury outboard for water skiing in 1963 about the time the new lab was finished

Laboratory for Limnology Leaders:
Art Hasler 1963 - 1978
Magnuson 1978 - 2001 
(Founding director of the Center for limnology
Jim Kitchel 2001 - 2009
Steve Carpenter - 2009 - Present (written in 2016)

Figure 120: Magnuson, Carpenter, Kitchel, Madison, 2009
(Center for Limnology Archives)

Art's successors 

Figure 121 Trout Lake Station Boulder Junction, 2016
(Aerial photo provided by Carol Warden)

Art also secured the initial Stage 1 funding in 1967 for the Limnology Station on Trout Lake, North of Minocqua WI. 

Stage 1 is the part of the building closest to the Lake.

Stage 2 (the new wing at right angles to stage 1) was built in 1985 and dedicated in 1989

Stage 3, built in 2012 (with the Juday conference room) is the section of the building farthest from the lake with the lighter roof. 

The Tom Frost House (adjacent but not shown) was built in 2009. It is used to house visiting scientists.

Figure 122: Trout Lake Station Boulder Junction, 2016
(Photo provided by Carol Warden)

The middle section is the new wing shown in Figure 173 when Art and UW Madison Chancellor, Donna Shalala, attended the dedication in 1989.

The section on the left with the entrance, houses the new conference room.

Figure 123: Old Labs at New Trout Lake Station, 2016
(Fritz Hasler Photo)

Susan Knight (Interim Director), ????, Fritz Hasler

Old Labs at the Trout Lake Station, Boulder Junction WI 2016

When the new lab was built in 1967 the old cabin labs, some built as early as 1926 were moved across the Lake on the ice to the new lab location. In 2016 the old labs are in great shape and look like they could have been built yesterday.

Figure 124:  Laboratory for Limnology, Madison, 1963
(Magnuson Viewgraph)

Figure 125: Art, Clifford Mortimer, new lab, Madison, 1963
(Center for Limnology Archives)

Figure 126: Art with Mortimer, New lab, Madison, 1963
(Center for Limnology Archives)

Mortimer was a visiting physical limnologist originally from the U.K. who had studied in Berlin Germany. He was later a professor at UW Milwaukee.

Figure 127: Art, Limnology Group, Madison, winter 1964
(Center for Limnology Archives)

Art far left.

Figure 128: Art, Henderson, Chipman, Madison, 1965
(Center for Limnology Archives)

Francis Henderson seated, Gerald Chipman standing on Lake Mendota
Tracking fish that have been released with inserted miniature transmitters.

Figure 129:  Art, Henderson, Madison, 1965
(Center for Limnology Archives)

Henderson holding a miniature ultrasonic transmitter to be stuffed into the mouth or anus of a fish (according to son Galen) so that it could be tracked (see previous figure)

Figure 130: Art, Zoology Faculty, Madison, 1966
(Center for Limnology Archives)

Art upper left in front of Birge Hall

Figure 131: Scherzl, Madison, 1965
(Arthur Hasler Photo)

As you would expect, Art was an excellent fisherman. Whether he was casting for walleye, dropping a line for perch or fly fishing on the Brule or the Yellowstone, he could do it all.

He was also an avid bird hunter, primarily for grouse and woodcock, but occasionally pheasant and quail. He had his favorite Browning double barrel over & under 20 gauge shotgun and always kept a German Shorthaired Pointer hunting dog, in the manner of the great Madison naturalist, Aldo Leopold. It wasn't a stretch for Art, the lover of Germany, to have a dog with German in it's name. The dogs always had German names. His first was Heidi, a solid chocolate, female which he bred several times for pure bred puppies, but his last was Scherzl, (in German little joke Art intended, but he later found our that with the L ending the word actually means the heel of a loaf of bread) the male shown above. He would also frequently practice with clay pigeons to improve his shooting.

Figure 132: Anneliese’s blessing dinner, Madison, 1968

(Fritz Hasler Photo)

A typical dinner, and number of guests, that Hanna served innumerable times, often to Art's professional guests, during her life.

In this image we see the assembled group and dinner table for the blessing of Anneliese (born December 4th 1967), the first child of Fritz and Mary Huebner Hasler and granddaughter of Art. Shown is the wonderful dining room with bay window of the Hasler 205 Lathrop Street house. The great round dinner table has a large Lazy Susan in the middle. A large turkey is carved in the foreground with mashed potatoes; on the Lazy Susan are homemade dinner rolls and the gravy for the mashed potatoes. 

Art is top row third from left and Hanna (far right) is in her element with a grand dinner for the blessing of their fourth grandchild, Anneliese.

Figure 133: Haslers, Thatchers, Madison,1968
(Arthur Hasler Photo)

Art & Hanna, their children, in-laws, and grandchildren shortly before Hanna died.

Top Row: Fritz, Mark, Karl, Christina, Bruce, Gilbert, Galen
Middle Row: Laurel, Hanna, Sabina, Arthur, Barbara, Sylvia
Front Row: Anneliese, Mary, Blaine


Lake Michigan Experiments: 
Imprinting and Homing of Salmon

Figure 134: Scholz meets Wisby, Madison, 1978
(Center for Limnology Archives cropped by AFH),

Hasler Retirement Reunion

Scholz upper left, Wisby lower right, Hasler center right.

Wisby began research on salmon homing with Art in 1949, determining that salmon have the capability of discriminating between different stream odors. In
 1951 he wrote an article proposing an experiment to prove that they use the capability for homing. Wisby conducted with Art the famous Washington State experiment where salmon noses were stuffed with cotton reported in the Wisby/Hasler article of 1954.

Scholz began with Art as an undergraduate assistant in 1968.  Scholz worked with Art for 15 years on 
Imprinting and Homing in Salmon in Lake Michigan, culminating in the comprehensive book (see below) on the subject. Some photographs of this effort from Scholz, figures from the book and Art's lecture slides with explanations and results are shown in the figures below.

Figure 135:  Hasler, Scholz, Imprinting & Homing, 1983
 (Cover of Hasler, Scholz, Olfactory Imprinting & Homing)

Working with Lake Michigan salmon, Scholz and Hasler conducted the experiment Wisby proposed in 1951 and wrapped up the loose ends with rigorous experiments proving that salmon imprint on a stream odor as smolts (fingerlings) and retain the memory of the odor for 1,5 to 5 years and use it to find their birth stream. The Hasler/Scholz collaboration culminated in the book Olfactory Imprinting and Homing in Salmon 1983 (see cover above).

Figure 136 Diagram from Hasler 1983 book with Scholz
(Hasler, Scholz, Olfactory Imprinting & Homing 1983)

This diagram shows where salmon were imprinted with two different chemicals (Morpholine and Phenethyl Alcohol) and and their return monitored in the two scented streams as well as most other possible spawning stream possibilities.

Figure 137 Diagram from Hasler 1983 book with Scholz
(Arthur Hasler Lecture Slide Carousel)

Color enlargement of the inset in the upper left corner of the previous figure.

Figure 138: Imprinting Experiment Results, 74 & 75 
(Arthur Hasler Lecture Slide Carousel)

Two Rivers/Little Manitowoc River, Wisconsin, 1974, 1975

The salmon imprinted with morpholine and phenethyl alcohol came back overwhelmenly to the streams where those chemicals were released compared to the controls that were not exposed to those chemicals.

Figure 139: Wild Rose Fish Hatchery, 1972
(Allan Scholz Photo)

Wild Rose Fish Hatchery near Wild Rose, not far from Waupaca in Central Wisconsin.

Raceways, Wild Rose Fish Hatchery where fish were held while imprinting them to MOR and PEA. Control fish were held in a third raceway

This is one of the two fish hatcheries where the salmon smolts were imprinted with either Morpholine or Phenethyl Alcohol. 

Figure 140: Scholz, Hirsch Clipping Fins, Wisconsin, 1972
(Allan Scholz Photo)

Scholz, Peter Hirsch (opposite) 2 other students at Wild Rose hatchery clipping fins of experimental fish, Wild Rose Fish Hatchery, 1972

Figure 141: Clipping fins of experimental fish

(Allan Scholz Photo)

Figure 142: Morpholine Fish, Little  Manitowoc R, 1974
(Allan Scholz Photo)

Note: Left pelvic fin is missing

"Two years later one of our fish identified by a left pelvic fin clip (this indicated it was a morpholine fish) that had returned to the Little Manitowoc River in 1974, where morpholine was being added to attract fish that had been exposed to morpholine during the smelt stage." (Scholz)

Figure 143: Crystal Springs Fish Hatchery
(Google Maps Satellite i.e. Aerial Photo View)

Actually this Crystal Springs Fish Hatchery is in Minnesota, I'm still looking for photographs of Crystal Springs Fish Hatchery near Antigo WI.

Crystal Springs is the other of the two fish hatcheries where the salmon smolts were imprinted with either Morpholine or Phenethyl Alcohol. 

Figure 144: Imprinting Hatcheries. Wisconsin. 1972
(Arthur Hasler Lecture Slide Carousel)

Locations of the Crystal Springs and Wild Rose Fish Hatcheries relative to the salmon release locations near Milwaukee and just south of Algoma (Manitowoc) Wisconsin.

Figure 145: Great Lakes Salmon Stocking
(Found on the Internet)

Figure 146: Scholz, Cooper, Gill Net,Two Rivers WI, 1974
(Allan Scholz Photo)

Scholz, Cooper, Setting Gill Net, East Twin River, Two Rivers WI, 1974

Figure 147: Scholz, Gill Net, Two Rivers WI, 1974
(Allan Scholz Photo)

Allan Scholz, Gill Net, East Twin River, Two Rivers WI, 1974

Figure 148: Salmon, Gill Net, Two Rivers WI, 1974
(Allan Scholz Photo)

 Salmon, Gill Net, East Twin River, Two Rivers WI, 1974

Figure 149: Scholz, Hughes, Goodman, E Twin River, 1974
(Allan Scholz Photo)

Allan Scholz, Kathy Hughes, and Cheryl Goodman catching salmon using electroshock, East Twin River, Two Rivers WI, just North of Manitowoc. Electroshock boat built by Frank Eustice and Scholz.

Figure 150: Hughes, Netting Salmon, E Twin River, 1974
(Allan Scholz Photo)

Salmon going into the live bin

Figure 151: Goodman, Hughes, Salmon, E Twin River, 1974
 (Allan Scholz Photo)

Cheryl Goodman punching the tail of a salmon with a paper punch while Kathy Hughes holds it. To prevent double counting in case of recapture.

Figure 152: Goodman, Art, Scholz, Electro-shocking, 1974 
(Arthur Hasler Lecture Slide Carousel)

Cheryl Goodman, Arthur Hasler, Allan Scholz, Electro-shock Fishing, East Twin River, Two Rivers Wisconsin 1974 

Using every technology to catch as many homing salmon as possible.

Figure 153: Scholz, DNR Biologist Crew, Algoma,1974
(Allan Scholz Photo)

Scholz and crew of Wisconsin DNR biologist checking for salmon returning to a DNR fish trap on the Ahnapee River, near Algoma, Wisconsin.

Figure 154: Kathy Hughes, Creel Census, Wisconsin, 1974
(Arthur Hasler Lecture Slide Carousel &  (Allan Scholz Photo))

Art liked to use attractive girls to do the creel censuses. The theory was that the fisherman was more likely to show her the over-the-limit fish he was hiding in his trunk.

Figure 155: Goodman, Creel survey, L. Manitowoc R., 1974
(Allan Scholz Photo)

"Cheryl Gosse Goodman doing a creel survey on Little Manitowac River. This was a girl data-blogger-escaped-comment-EndFragmentwho was an undergrad hourly assistant who took a semester off to help us perform this work. She later took a job as a fish biologist with the Wisconsin DNR" (Scholz) 

They used every method possible including interviewing fishermen, gill netting and electrofishing to examine every returning salmon possible for the markings done when imprinting.

Figure 156: Art, L. Michigan Salmon, Strawberry Creek, 1974
(Center for Limnology Archives)

 "Art Hasler in October 1974 being interviewed on the successful homing of Pacific salmon at Strawberry Creek on the Wisconsin shore of Lake Michigan. 

This was a part of a Sea Grant funded project testing whether young salmon hatched and imprinted to two artificial chemicals returned to the correct stream with the imprinted odor being released into it. "  (John Magnuson)

Note the microphone held in the hand of the interviewer mostly out of view on the left.

Figure 157: Placing Ultrasonic Transmitter, Wisconsin,1972
(Arthur Hasler Lecture Slide Carousel)

Placing an ultra-sonic transmitter into the mouth of a salmon

Figure 158: Scholz, Salmon Tracking, Wisconsin, 1972
(Arthur Hasler Lecture Slide Carousel)

Ultra-sonic Salmon Tracking at Oak Creek in South Milwaukee.

Figure 159: Scholz, Tracking, S. Milwaukee, 1972
(Allan Scholz Photo)

Scholz ultra-sonic tracking in a snow storm, Oak Creek, South Milwaukee Wisconsin, Fall 1972

Figure 160: Scholz, Tracking, S. Milwaukee, 1972
(Allan Scholz Photo)

Scholz ultra-sonic tracking Salmon, Oak Creek, South Milwaukee Wisconsin, Fall 1972

Figure 161: SalmonTracking Results, Wisconsin. 1972
(Arthur Hasler Lecture Slide Carousel)

Ultrasonic SalmonTracking Results 1)

 Imprinted salmon (a.) would loiter in the decoy area where the imprinted chemical was released. However the controls (non imprinted salmon) would pass by the decoy without hesitation.

Figure 162: Salmon Tracking Results, Wisconsin_ 1972
(Hasler/Scholz Book 1982)

Ultrasonic SalmonTracking Results

 With no chemical release (A.), or with fish imprinted with phenethyl alcohol the salmon would not even swim up-stream. When morpholine was released (B. & C) the morpholine imprinted salmon would swim upstream along an edge of the morpholine plume. As Art would say, much like a hound tracking a fox: crossing back & forth over the odor trail, not staying in the middle of the trail where the nostrils would become oversaturated with the smell.

Figure 163: Fish, Electrode in the olfactory bulb of its brain
(Allan Scholz Photo)

Fish with an EEG electrode placed in the olfactory bulb of its brain

Figure 164: Testing a fish’s EEG response to various odors
(Hasler/Scholz Book 1982)

Amazing that you can observe in the brain of an adult salmon the response to the chemical it was imprinted to as a smolt (fingerling)

"Testing a fish’s EEG response to various odors; a) response of adult fish that was exposed to morpholine as a smolt to morpholine; b) response of same fish to phenylethyl alcohol (PEA); C) response of same fish to Oak Creek water that was scented with morpholine; d) response of same fish to Oak Creek water that did not contain morpholine [Note: c and d were interesting because in this experiment Oak Creek was scented with morpholine below a dam located about two miles up the river. We added the odor below the dam to insure better mixing of the odor into the creek and because we knew the volume of water flowing over it, so we could add morpholine to it to achieve a known concentration of 5 X 10 -5 mg/L morpholine in the creek. Later we collected water samples below (contained morpholine) and above (did not contain morpholine as a control) the dam and tested the response of adult fish that had been exposed to morpholine as a smolt to each sample. e) morpholine exposed fish response to nBhydroxyethyl – morpholine. f) response of a fish exposed to PEA as a smolt to morpholine."

Art's daughter Sylvia writes: 

"We lived in Milwaukee WI at the time the salmon were being tempted into the Oak Creek river in southern WI with the chemical morpholine to which they had been imprinted.    My father Arthur Hasler invited me to bring my 5 children over to the Oak Creek High school where they, his researchers, were set up in the science lab of the school and showed us a salmon with his brain discected open as they demonstrated the reaction in the brain of the living salmon when they would pour normal Lake Michigan water over the brain and then the frantic neural activity reacting to the change to morphelin laden water transferred to a screen on which the electrical activity was demonstrated.  It was very exciting to see this dramatic change to the response of the salmon to the water to which it had been imprinted to (as a smolt) to return to it's home stream at the proper development age. These salmon were now pouring into the mouth of the Oak Creek as it entered Lake Michigan. The scientists graciously gave us a female salmon to take home with us which our 3 year old proceeded to play with in the sink and bathtub as we debated about how to preserve the plentiful  roe for caviar." ++++++++++++++++++++++++++++++++++++++++++++++

Note: Part II of the book Olfactory Imprinting and Homing of Salmon entitled:

Hormonal Regulation of Smolt Transformation and Olfactory imprinting in Salmon 

is not covered in this blog post.



Figure 165: Art, Peter Hirsch, Madison, 1976
(Center for Limnology Archives)

Conditioning the hearts of fish to electric shock.
Using a method for determining whether a fish can detect an oder

Figure 166: Lake Como Italy, 1977
(Center for Limnology Archives)

Art and his Olympus Pen F half frame camera,

Many of the pictures in this blog were taken by Art himself, but this is the only one we have of of him with a camera.

Earlier he had numerous Leica cameras, with which he took thousands of color slides starting about 1940.

Figure 167: Leica 111c, Leica M3, and Pen F cameras.
(Assembled from Internet photos by AFH)

Art liked compact, mostly German 35 mm cameras. He started shooting Kodachrome color slides about 1940 with the Leica 111c and upgraded in the 1950s to a Leica M3. He always carried and used a light meter.  He took his photography very seriously.

When his son Fritz started using the even more compact half frame Olympus Pen F with built-in light meter in the 1960s, Art later followed suit (see figure 129 above), For a period, Art also had a larger format Rolleiflex twin lens view camera.

Figure 168: Art, Madison, 1977
(Center for Limnology Archives)

Limnology Lab Leader, in the Corner Office.



Figure 169: Art, Teaching Class, Madison, 1977
(Magnuson Viewgraph)

Figure 170: Arthur Hasler Lecturing, Madison, 1985
(Center for Limnology Archives)

In addition to all his other activities, Art was an excellent teacher. We found his Kodak Slide Carousel that he used for his standard lecture when traveling. It included several slides (Figures 132 - 145 below) that showed the process and results of the Salmon imprinting and homing experiments with Scholz on Lake Michigan in the 1970s. The next Figures show some the slides that came from his lifetime of experience and world wide travels.

Figure 171: Joe Merrill, Salmon, Alaska, 1965
(Arthur Hasler Lecture Slide Carousel)

Art's daughter Sylvia writes:

"(Joe Meril)  rented a stretch of beach on the Kachemak Beach on the Kenai Peninsula in Alaska for the summer. He and other fishermen would set a line of baited hooks on a string and then would go out each day to take the fish off the line and a truck would come by to pick up their catch each day.  He made $ 25,000 a summer each year from 1964, 65, and 66.  It put them thru graduate school in Madison Wis at the unit of WI. She said that was the only time in their married life when she felt like she had more money than  she could spend.  He later taught Accounting at  Univ of Alaska in Anchorage and several other places, Logan and Cedar City Utah"

Figure 172: Salmon Spawning Cartoon
(Arthur Hasler Lecture Slide Carousel)

"Well, that's about it, baby! We spawn right here or we don't spawn at all"

He was usually very serious, but he had a sense of humor.

Figure 173: Lake Baikal, Russia, Map
(Arthur Hasler Lecture Slide Carousel)

He could lecture on all the "Great" Lakes of the world and had visited all of them.

Figure 174: Olfactory System of a Bony Fish, von Frisch
(Arthur Hasler Lecture Slide Carousel)

A slide of a von Frisch illustration. It was one of his favorites and must have been in his collection since the 40s. It was one on the Olfactory System of a (Knochenfisch) Bony Fish. It was from one of the articles by von Frisch that led him to go to visit him in 1945 in Austria and begin a life-long friendship.

Figure 175: Salmon Jumping
(Arthur Hasler Lecture Slide Carousel)

The man who figured out how salmon home, had to have at least one picture of salmon jumping a waterfall. He also had a few pictures of salmon spawning. None as good the ones I have in Figures 69 and 70. 


Art  is elected to the National Academy of Sciences
in 1969 and receives many other honors throughout his career

Figure 176: Art, National Academy of Sciences, 1990
(Arthur Hasler Photo)

Image with the Einstein statue, supplied by Hatheway, I'm looking for a higher resolution version

Figure 177: Art, Einstein Statue, Washington DC
(Found on the Internet)

 Photo taken at the National Academy of Sciences Headquarters in Washington DC

Figure 178: Arthur, Hatheway, Mary, Marta, Annapolis, 1977

(Fritz Hasler Photo)

After the death of his first wife Hanna Prusse in 1969 Art married Hatheway Minton (see above) in 1971.

Art was elected as a member of the National Academy of Sciences in 1969. In American scientific circles this is an honor short only of winning the Nobel prize. 

In this photo Art and Hatheway have traveled from Wisconsin to Washington DC to attend the annual meeting of the Academy. He is visiting his eldest son Fritz, his daughter-in-law Mary (see above) and grand children (see Marta above) The photo is taken at the Annapolis City Dock.


Art Retires 1978 at big Retirement Party Held in His Honor

Figure 179: Art, Small group retirement, Madison, 1978
(Center for Limnology Archives)

Back Row: Bill Helm, Warren Wisby, Ross Horrall, Bob Ragotzkie, Ken Johns

Front Row: Art, John Bardach, Ralph Kursaal

Figure 180: Art, Group Retirement, Madison, 1978
(Center for Limnology Archives)

Art middle row center, Ragotzkie upper left, Wisby center front, Horrall, 1st row 3rd from left. Gene Likens 1st Row 2nd from right. Bill Schmitz 2nd Row 4th from left.

1st Row: Ken John, Jim Gammon, Ross Horrall, Bill Helm, Joe Koonze, Warren Wisby, Mike Park, Paul Sager, Gene Likens, Ray Stross,

2nd Row: Libby Jones, Peter Hirsch, Ralph Nursall, Bill Schmitz, Ray White, Art, Tom Wissing, Jon Cooper, Dan Faber, Jim Jager, Clyde Voightlander, Kenton Stewart

3rd Row: Bob Ragotzkie, Peter Johnson, Nick Lenz, Clarence McNabb, Henry Eichorn, Al Scholz, George Gallepp, John Bardach, Bob Hunt, Arne Salli, Mits Teraguchi, Phil Doepke, Gary Hergenrader, Horst Schwassman, Jim Bruins.

Figure 181: John Magnuson, Madison, 1978
(Center for Limnology Archives)

Art retired at age 70 and turned over the reins and Leadership of the Laboratory for Limnology to his colleague John Magnuson. Magnuson served as Leader from 1978 until 1982 when he founded the Center for Limnology and became its first director. He retired in 2001

Magnuson was extremely helpful in collecting the photographs (along with my brother Galen) and pointing me to literature that I should read in order to put this blog post together.

Figure 182: Art, Hatheway, Madison, 1978
(John Magnuson Photo)

Art and his second wife Hatheway Minton at a dinner at John Magnuson's house.

Figure 183: Art, Madison, about 1980
(Center for Limnology Archives)

Art, Emeritus in his office 

According to his daughter Sylvia: still bringing in million dollar grants.

Figure 184: Tom Frost, Little Rock Lake, 1991
(John Magnuson Photo)

Double Neoprene barrier for whole lake experiment where sulphuric acid was added to one side and the other was left as a control.

Figure 185: Sulphuric Acid, Little Rock Lake, 1991
(Center for Limnology Archives)

Preparing Sulphuric Acid for insertion into lake

This was the inverse of the experiments with Peter & Paul Lakes in the 1950s where the acidity of the lake was decreased by adding lime.

In this case the acidity of the lake was increased by adding many gallons of sulphuric acid  to the water and comparing the changes in the lake to the part of the lake on the opposite side of the neoprene barrier.

Just to prove that UW limnologists, in the tradition of Arthur Hasler, weren't afraid to mess with mother nature in order to advance the scientific knowledge of how lakes function.

Figure 186: Art, Madison, 1984
(Center for Limnology Archives)

Looking at Lake Mendota Plankton

Figure 187: Art, Fritz von Siegfried, Madison, 1984
(Center for Limnology Archives)

von Siegfried was a famous nature photographer and cinematographer.

This photograph may well have been taken in Art's beloved UW Arboretum in Madison. Art was a very active member of the Arboretum Board for some 30 years and worked tirelessly to protect it, expand it and make it available for scientific studies.

Figure 188: Art calling annual spring cleanup, Madison, 1986
(Center for Limnology Archives)

Figure 189: Magnuson, Art, Clean-up Crew, Madison, 1986
(Center for Limnology Archives)

In front of the Limnology Lab: Art was a stickler for cleaning up trash where ever he went.
From his daughter Sylvia: Art was walking along the lake with four distinguished scientists talking about their work. According to his usual habit Art is picking up trash as he goes. When they arrive at the lab, all five of them have both hands full of trash.

Figure 190: Beckel Quote, 1987
(John Magnuson Viewgraph)

As his son and the only one of his boys to get a PhD, in researching this blog post, I am continually astonished how many balls my dad kept in the air at one time.

I purposely went to a government Lab (to NASA Goddard Space Flight Center) instead of a University, because I had too many professors that were bad teachers from spending too much time with their research and publishing. I didn't think I could pursue a career in research and be a good teacher at the same time

Furthermore, the number of relationships he maintained with other scientists in addition to doing his own research and mentoring his students is amazing.

Figure 191: Art, President Pearson, Oxford Ohio, 1988
(UW Madison Archive)

Art, Miami University President Pearson, Oxford Ohio, 1988

Art receiving the honorary Doctorate of Science degree from Miami University presented by the president of the University

Art was elected a member of the National Academy of Sciences in 1969. In American scientific circles this is an honor short only of winning the Nobel prize. He also received numerous other awards and honorary degrees from other Universities like the one shown above

Figure 192: Art,  Shalala, Trout Lake, 1989
(Center for Limnology Archives)

Dedication of the new wing of the Trout Lake Station 07June1989

Art loved to kiss the hand of a women he was meeting.

Donna Shalala: The Honorable United States Secretary of Health and Human Services from 1993-2001
Chancellor of the University of Wisconsin-Madison from 1988 to 1993

Figure 193: Art, Douglass Hill, Madison, 1990
(Center for Limnology Archives)

Douglass Hill with his quartet after his performance of his French Horn composition at an anniversary party for ADH. Hill followed Art as a French Horn Player with the Madison Civic Symphony and is now a retired Professor of the University of Wisconsin School of Music.

Figure 194: Hatheway, John Bardach, Art, Madison, 1990
(Center for Limnology Archives)

Art Receiving the music for a  French Horn composition by Douglass Hill after his performance at an anniversary party for ADH.

Arthur married Hatheway Minton, July 24, 1971 in Madison Wisconsin and was married to her for 30 years until his death in 2001.

Hatheway was an immeasurable aid to Art during his later career and during his declining years.

He was married to his first wife Hanna for 37 years

Figure 195: Art, Receives AIFRB Award, Madison, 1993
(Center for Limnology Archives)

Art Receives AIFRB (American Institute of Fishery Research Biologists) award.

Top row left to right:  Kitchel, Art, AIFRB President, Magnuson, Carpenter 

Seated: Linda Holthaus, Hatheway.

Art was also awarded the Navmann-Thienemann Medal from the International Association of Theoretical and Applied Limnology, the highest international award in Limnology in 1992

Figure 196: Art, Madison, about 1995
(Center for Limnology Archives)

Figure 197: Art, Europe, about 1998
(Center for Limnology Archives)

Art with monocular telescope and hand lens around his neck.

In his declining years, Art suffered from macular degeneration of his retinas. He was essentially totally blind except for his peripheral vision which was only a blur. He carried monocular telescope around his neck, but I looked through it once and I'm not sure it helped him as he didn't even have the focus set properly.

Figure 198: Arthur, Sylvia, Limnology Lab, Madison, 1998
(Fritz Hasler Photo)

16Sept1968: Arthur with daughter, eldest child Sylvia, in front of the Limnology Lab.

Figure 199: Arthur, Limnology Garden, Madison, 1998
(Fritz Hasler Photo)

September1998: The Limnology Garden honoring Arthur Davis Hasler

Figure 200: Art, Madison, 1999
(UW-Madison Archives)

Dressed for Madison's occasional bitter cold,

Figure 201: Art, Fritz, Matt, Trout Lake, 2000

(Fritz Hasler Photo)

20July2000: Back at Trout Lake once again over 67 years - Three generations of Eagles

Arthur Davis Hasler, son Arthur Frederick (Fritz) Hasler, and grandson, Matthew Frederick Hasler. This may be the first time Art had worn his Eagle Sash in 77 years (see Figure 3, the 1924 photo when he became an Eagle at 16).

Art would be dead seven months later.


As recalled in the biographical synopsis at the beginning of this blog post:

 Art survived four kinds of cancer, two surgeries, and chemo therapy. He wore a colostomy bag for the last 20 years of his life which he handled with amazing competence and good cheer. 

However, the last 10 years, particularly the last two years of his life were very sad. For the last 10 years of his life, he suffered from macular degeneration of his retinas. He was essentially totally blind except for his peripheral vision which was only a blur. He carried monocular telescope around his neck, but I looked through it once and I'm not sure it helped him as he didn't even have the focus set properly. The last two years he suffered from dementia which got progressively worse.  Hatheway was a true saint, to guide and help him through his last years.

I won't do this justice, but near the end of his life, I remember him saying something like this to Hatheway, his wife of 30 years: "I know you, I know your name, it's right on the tip of my tongue, help me with it, please help me"

Even with dementia at this level. dad would repeat over and over his favorite saying about the German language (see the caption to Figure 6 1928) and his favorite German Poem:

Er ist’s
Frühling läßt sein blaues Band
Wieder flattern durch die Lüfte;
Süße, wohlbekannte Düfte
Streifen ahnungsvoll das Land.
Veilchen träumen schon,
Wollen balde kommen.
– Horch, von fern ein leiser Harfenton!
Frühling, ja du bist's!
Dich hab ich vernommen!
Eduard Mörike (1804 – 1875)
Copy the poem into Google Translate for a rough translation into English

Figure 202: Eagle Sashes, Trout Lake, 2000.

(Fritz Hasler Photo)

Three generations of Eagle Sashes. Art: 1924, Fritz 1961, Matt 2000 (in Figure 3 you can see Art wearing the Eagle pin in 1924)

Figure 203: Art's Funeral, Madison, 2001

(Fritz Hasler Photos)

20March2001: Hatheway, son Karl, Jim Kitchel, LDS Chapel, Madison

Figure 204: Some of Art's Decedents. Stewart Falls, 2002
(Fritz Hasler Photo)

Some of Art's Decedents Stewart Falls, Utah, 2002

The scent that changed the world, 

Art's grandchildren and great grandchildren at Stewart Falls on Mt Timpanogos near Provo Utah. This is the Falls that Art smelled before it came into sight that triggered his recollection of "childhood chums" and gave him the insight to find out if sense of smell was the tool that salmon use to find their home stream. He was taking his young sons on a hike about 1950.

It was during a vacation to Utah that an incident occurred that caused Hasler to incorporate the imprinting concept with the problem of homing in salmonWe had driven across the sage country and high desert from Madison, Wisconsin to my parental home in Provo, Utah . As I hiked along a mountain trail in the Wasatch Range of the Rocky Mountains where I grew up, my reflections were interrupted by wonderful scents that I had not smelled since I was a boy. Climbing up toward the alpine zone of Mt. Timpanogos, I had approached a waterfall which was completely obstructed from view by a cliff; yet, when a cool breeze bearing the fragrance of mosses and columbine swept around the rocky abutment, the details of the waterfall and its setting in the face of the mountain suddenly leapt into my mind’s eye. In fact, so impressive was this odor that it evoked a flood of memories of boyhood chums and deeds long since vanished from conscious memory.  The association was so strong that I immediately applied it to the problem of salmon homing.  The connection caused me to formulate the hypothesis that each stream contains a particular bouquet of fragrances to which salmon become imprinted before emigrating to the ocean, and which they subsequently use as a cue for identifying their natal tributary upon their return from the sea. I envisioned that the soil and vegetation of each drainage basin would impart a distinctive odor to the water, thereby providing the salmon with a unique cue for homing” (A. D. Hasler, in Hasler and Scholz 1983).


Art's Legacy

The students who carried on Art's scientific traditions are Art's greatest legacy

Figure 205: AD Hasler Genealogy Tree, Madison, 1978
(Fritz Hasler Photo of Kandis Elliot Drawing)

Artistic genealogy tree drawn by Kandis Elliot showing all of Art's Masters and PhD students.

52 doctoral students and 43 masters students received their degrees under his supervision

A complete list of Art's MS and PhD student and their thesis topics can be found in the Appendix of Breaking New Waters, a Special Issue of the Transactions of the Wisconsin Academy of Arts and Letters.

Figure 206: Black & White Genealogy Tree, Madison, 1978
(Fritz Hasler Photo of Kandis Elliot Drawing)

If you zoom in on this Black & White tree you may be able to read the text (see CU below)(.

Figure 207: Section of Genealogy Tree, Madison, 1978
(Fritz Hasler Photo of Kandis Elliot Drawing)

A section of the genealogy tree with his first PhD students.

His first student was Jay Andrews who got his PhD in 1946. He had three students get degrees in 1947 and his last students Peter Hersch PhD, Eric Olsen PhD and Alan Scholz MS got their degrees 30 years later in 1977.

Some notable students from this history and my memory were:

John Bardoch PhD 1949, John Nees PhD 1949, Warren Wisby PhD 1952, Bob Ragazkie PhD 1953, Kenneth John PhD 1954, Bill Helm PhD 1958, Bill Schmitz PhD 1958, Ross Horrall PhD 1961, Gene Likens PhD 1962, Francis Henderson PhD 1963, Don McNaught PhD 1965, Peter Hirsch PhD 1977, Allan Scholz, 1977

Figure 208: Center for Limnology, Faculty, 2009
(Center for Limnology Archives)

Current Faculty from left to right: Paul Hanson (Research Professor), Steve Carpenter (Director), Jake Vander Zanden (Professor), Emily Stanley (Professor), Pete McIntyre (Assistant Professor), John Magnuson (Emeritus Director)

The Laboratory of Limnology Facility and the scientists that work there now is an equally important legacy of Arthur Davis Hasler

Figure 209: Lab Naming, Madison. 2006
(Fritz Hasler Photo)

08May2006:, Widow Hathaway, and children, Fritz Sylvia, Karl, Mark, Galen

Arthur's widow, his second wife, Hatheway, and five of his six children along with 100s attend the naming ceremony for the A. D Hasler Laboratory of Limnology which he built.

His professional legacy is the ongoing labs in Madison and Trout Lake and the nearly 95 PhD and Master students that received their degrees under his supervision. His 200 pier reviewed papers and seven books have inspired countless others to continue his work.

Figure 210: Lab Naming, Madison. 2006
(Fritz Hasler Photo)

Art's decedents at the lab naming

Figure 211: Wiley, Lab Naming, Madison. 2006

(Fritz Hasler Photo)

University Chancellor, John Wiley, speaking at the Lab naming, May 2006

Figure 212: Audience, Lab Naming, Madison. 2006

(Fritz Hasler Photo)

Figure 213: Lab Naming, Madison. 2006

(Fritz Hasler Photo)

Lab Director, Jim Kitchell and the extended Hasler family, on a boat and pier in front of Limnology Laboratory

Figure 214: Lab Naming, Madison. 2006

(Fritz Hasler Photo)

Audience for Lab Naming Ceremony and Luncheon at the UW Pyle Center

Figure 215: Salmon, Lab Naming, Madison, 2006
(Fritz Hasler Photo)

Salmon with noses stuffed with cotton, served at the naming ceremony in May of 2006.

Recalling Art's genius to capture salmon above a fork in a river, take them below the fork, stuff the noses of half of them and release them, capture them again above the fork to prove that only the salmon without stuffed noses could tell which way led to the stream of their birth.
Art's family is another big part of his legacy

Without them this photo biography would not have been possible

Art's second Wife Hatheway Minton and all of Art's six children are alive and in relatively good health as this is written 15 years after his death in 2016 .

As this is written in 2016, he has 18 grandchildren, 59 great grandchildren, and 3 great great grandchildren.

Figure 216: Art's Children, Three Lakes WI, 2012
(Fritz Hasler Photo)

Art & Hanna’s six children: Sylvia, Fritz, Mark, Galen, Bruce, Karl

Sylvia was the keeper of Art's letters from Germany in 1945, Fritz is the author of this blog post, Galen was instrumental in finding many of the photographs presented here, Galen Mark, Bruce, Karl and Sylvia were very helpful with encouragement, suggestions, and corrections to this piece.


Art's Portraits Through the Years


1916 Age 8:  Provo  

  1924 Age 16: Eagle Scout


1925 Age 17: High School Junior 


 1927 Age 19: BYU Freshman

1929 Age 21: Missionary Passport Photo

1932 Age 24: BYU Senior

1933 Age 25: UW Grad Student 

1933 Working with Birge & Juday: Trout Lake

1937 Age 29: Crater Lake Ranger  

1940 Age 32: UW Instructor

1943 Age 36: Assistant Professor 

1945 Age 37: Associate Professor



1945 Major Hasler: US Army   

 1950 Age 42: Full Professor



1953  Age 45: Portrait  

1955 Age 47: Fulbright Professor to Munich
(Arthur Hasler Photo)


1956 UW Photo Services Portrait 

1956 Casual Portrait



1958 UW Photo Services Portrait 


1958 SLC: On The Move
(Arthur Hasler Photo)

1958 Home for Christmas

1965 Age 57: Limnology Laboratory Leader     

1977 Age 69: Lab Leader Near Retirement

1977 Lab Leader Near Retirement

1980 Age 72: Emeritus in his Office

 1986 Age 78: Conch Cleanup Call


1995 Portrait

1995 A Winter day in Madison




1998 with Monocular   

1998 with Monocular                                       

2000 Age 92: Winter in Madison shortly before his death


Gene Likens biography of Arthur D Hasler

January 5, 1908–March 23, 2001
ARTHUR DAVIS HASLER did pioneering limnological research across a broad spectrum of ecological subdisciplines from ecophysiology and behavior of fish to experimental manipulation of entire lake ecosystems. His work on the mechanisms whereby salmon find their way back from ocean feeding areas to home streams for spawning, for which he was best known, was not only brilliant and innovative but also provided a framework for management of these important fisheries throughout the world.
Hasler was born in Lehi, Utah, the third of four sons of Mormon parents who also had one daughter, Walter Thalmann Hasler, a physician, and Ada Broomhead Hasler. His Mormon background played a significant and important role throughout his life, particularly regarding his active role in public service. He was among those who strongly advocated for acceptance of African-American membership in the Mormon Church.
He married Hanna Prusse in 1932, and they had six children: Sylvia, A. Frederick, Bruce, Galen, Mark, and Karl. Hanna was a trained vocalist (soprano) and music was a large part of the family’s activities. Hasler’s passions went far beyond science. His love of music and poetry was legend among his students and colleagues. He recited the works of Mörike, Heine, or Goethe at every opportunity and played
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the horn (waldhorn) for some 30 years in the University of Wisconsin Symphony and the Madison Civic Symphony. He frequently greeted a woman with a kiss to her hand. On long road trips to research sites and scientific meetings it was not uncommon for Hanna to break out the songbooks, pass them out in the car, and lead everyone in singing. In those days a major professor and graduate students often took long trips together by car to field sites and professional meetings. Hanna died in 1969. In 1971 Hasler married Hatheway Minton Brooks, who shared his love for a healthy environment, and with her own six children forged a close and loving extended family. Hasler had 32 grandchildren and 17 great-grandchildren.
He received a B.A. degree, majoring in zoology, from Brigham Young University in 1932 and a Ph.D. degree in zoology and physiology from the University of Wisconsin-Madison in 1937. He was awarded honorary doctor of science degrees from the Memorial University of Newfoundland in 1967 and Miami University, Oxford, Ohio, in 1988.
Hasler had interrupted his schooling at Brigham Young University in the late 1920s to serve a three-year mission to Germany for the Church of Jesus Christ of Latter Day Saints. It was during this time that his love for the German language began.
After working for the U.S. Fish and Wildlife Service as an aquatic biologist on the Chesapeake Bay during 1935–37, he and Hanna moved to Madison where he completed his Ph.D. in 1937 at the University of Wisconsin, under the supervision of well-known limnologist Chancey Juday. He was hired there as an instructor of zoology in 1937 and promoted to assistant professor in 1941. After serving with the U.S. Strategic Bombing Survey in Germany in 1945, he returned to the university in 1945 as associate professor of zoology and was promoted to full professor in 1948, and
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served in that capacity until he retired in 1978. During that time 52 doctoral students and 43 masters students received degrees under his supervision.
Hasler actively published in the peer-reviewed literature for almost 50 years from 1935 to 1984. He authored, co-authored, edited, or contributed to 7 books and over 200 scientific publications.
For more than a hundred years the University of Wisconsin-Madison has been an international center for limnology. Started by Edward A.Birge and Chancey Juday in the late 1800s, the Wisconsin School of Limnology was continued, strengthened, and enlarged by Hasler from 1946 to 1978. He supervised a large, active, and diverse limnology program conducted in several scattered and some rather Spartan structures on campus, known affectionately as the Lake Lab. In 1963 he became director of the Laboratory for Limnology coincident with the construction of a new and proper Limnological Laboratory on the shoreline of Lake Mendota. He fought aggressively and successfully with the faculty and administration of the university against the construction of a 600-car parking lot on the site and extending into Lake Mendota. His final plea at the faculty hearing was a quote from St. Mark: “Go thy way and sin no more.”
Hasler was one of the preeminent ecologists of the twentieth century. When he was elected to the National Academy of Sciences in 1969 only two other ecologists (G.E. Hutchinson and C.L.Hubbs) had ever received this prestigious honor. Hasler was a Fulbright research scholar in Germany in 1954–55 and a Fulbright visiting professor at the University of Helsinki in 1963. He was elected to the Societas Scientiarum Fennica in 1965, the American Academy of Arts and Sciences in 1972, the Royal Netherlands Academy of Science in 1976, and the Wisconsin Academy of Sciences, Arts, and Letters in 1988. He received 10 distinguished
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scientist awards, including the Award of Excellence from the American Fisheries Society in 1977, the Distinguished Service Award from the American Institute of Biological Sciences in 1980, and possibly most significantly, the Citizen of the Year Award from the Mendota-Monona Lake Property Owners Association in 1987.
An important measure of his influence in professional biology was his service as president of the American Society of Limnology and Oceanography (in 1951), the Ecological Society of America (in 1961), the International Association for Ecology (1967–74), and the American Society of Zoologists (in 1971). Hasler also was the founding director of the Institute of Ecology (1971–74). He was awarded the Naumann-Thienemann Medal from the International Association of Theoretical and Applied Limnology, the highest international award in limnology, in 1992. He was an exchange scholar for the National Academy of Sciences in China in 1983 and in the Soviet Union in 1986.
With very broad interests and expertise he could equally well have carried the scientific descriptor of limnologist, ecologist, fishery biologist, zoologist, and conservationist. He conducted research and informed public policy in all of these disciplines.
Hasler is best known for his research on salmon olfactory imprinting, a powerful and ingrained sense of smell that enables these fish to return to the exact stream of their birth for spawning after traveling thousands of kilometers in the ocean. He often told the story about the genesis of this discovery when he was vacationing in the Wasatch Range of the Rocky Mountains of Utah, where he had spent much time as a boy. Hiking up a mountain, yet out of sight of his favorite waterfall, he suddenly had what he called a “déjà senti” experience, “as a cool breeze, bearing the fragrance of mosses and columbine, swept around
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the rocky abutment, the details of this waterfall and its setting on the face of the mountain suddenly leapt into my mind’s eye” (1966, p. 65). Among other things these smells reminded him of childhood memories and of home. If smells could trigger such memories in a human, they must be at least as evocative for salmon, Hasler reasoned. This revelation led to a rich and productive series of experiments and field trials on olfactory and solar orientation in fishes.
Hasler’s pioneering research using manipulation of entire lake ecosystems provided a powerful new tool for ecology. Following the early lead of his major professor, Juday, who had added fertilizer to lakes to increase fish production, Hasler greatly developed and expanded this new experimental approach for studying large ecosystems (lakes) within their natural settings. He recognized early that entire ecosystems were just too complex to study piecemeal or only in the laboratory. His first efforts were focused on trying to enhance the productivity of fish in the thousands of acidic brown-water lakes in the upper midwestern United States. The brown staining by dissolved organic matter in these lakes prevented light penetration and thereby reduced productivity of aquatic plants at the base of the food web. In 1947, finely ground hydrated lime (calcium and magnesium hydroxide) was added to an acid, brown-water lake in Langlade County, Wisconsin, to determine whether the water could be cleared and the depth of the trophogenic zone increased. This experiment was aborted. Then, in 1950 hydrated lime was added to two small lakes in Chippewa County, Wisconsin, and resulted in remarkable alkalinization and increased transparency of the water (1951). The stage was set for a rigorous whole-lake experimental manipulation, so he found two lakes on the northern Wisconsin-Michigan border, located on property of the University of Notre Dame, that were connected and together were
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shaped like spectacles or an hourglass. He obtained permission to bulldoze an earthen dam across the narrow constriction between these two lakes in 1951 and thus formed the now famous setting for whole-lake experiments by creating two separate lakes, Peter and Paul. Subsequently, Peter Lake was treated with hydrated lime to flocculate and precipitate the dissolved organic carbon in these humic brown-stained lakes, while Paul Lake was maintained as an untreated reference in this experimental manipulation (Stross and Hasler, 1960).
Other lakes were artificially circulated using compressed air to reduce ice cover and prevent winterkill of fish, experimentally manipulated with additions of hydrogen peroxide to reduce color, manipulated by additions of hydrated NH4 through rather primitive aeration systems on the bottom of lakes to increase productivity, and labeled with radioactive tracers to study water circulation and biological transport of nutrients from deepwater to surrounding landscapes. These field experiments in whole-lake ecosystems had varied levels of success, but the overall approach was innovative and powerful. Inspired by this model for the study of complex natural ecosystems, W.E.Johnson, one of Hasler’s Ph.D. students, and J.R.Vallentyne designed an experimental lakes area in Ontario, Canada, that used whole-lake manipulation very successfully in studies of lake eutrophication, acidification, and toxification by heavy metals. Likewise, F.H.Bormann, R.S.Pierce, N.M.Johnson, and the author (another Ph.D. student of Hasler’s) adopted an experimental approach in studies of watershed ecosystems in the Hubbard Brook Valley of New Hampshire. This small watershed approach—in association with a nutrient flux and cycling model and entire watershed manipulations—helped establish fundamental understanding of northern hardwood forest ecosystems.
Hasler always freely acknowledged the role of travel and
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his many professional colleagues, students, and visitors in expanding his reach in scientific inquiry and influence. He insisted that graduate students in residence meet with and discuss their research with each visiting scientist. Because of the international stature of his program, there was a constant flow of visitors to the Lake Lab. Hasler’s contemporaries were the noted animal behaviorists Nobelist Karl von Frisch and Konrad Lorenz; Wilhelm Einsele, limnologist and chemist; and G.Evelyn Hutchinson, limnologist and ecologist. He considered them scientific heroes.
Hasler made it a point to provide not just academic training for students but personal advice as well. He usually had a large number of students under his supervision, but he took special interest in each of us. His achievements, career, and style were an inspiration for us, and he invested much time promoting his students.
The National Science Foundation was just beginning to fund science shortly after Hasler started his research career at the university. He successfully obtained financial support for his research from the Atomic Energy Commission, Office of Naval Research, and of special significance, wealthy landowners in northern Wisconsin. Several research projects were supported for decades by and on the properties of these philanthropists. He was able to convince these landholders of the practical importance of this research, and thus of its benefit to them.
Not only was Hasler a preeminent scientist but he was also a preeminent statesman of science. Constantly working to enhance organizations, networks, and teams to promote the betterment of ecology and the conservation of natural resources, he had few peers, and his efforts have provided a continuing legacy. Late in his career he tried to initiate a “Salmon for Peace” project, which attempted to bring together the governments of Russia and China to restore and manage
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the salmon population in the Amur River, which had been depleted because of overfishing. Even though this effort was unsuccessful, it clearly demonstrated his desire to apply ecological understanding to practical problems.
Hasler played a key role in developing and promoting the fact that land-water interactions are important for what occurs in lakes, such as variable water quality. His early classic paper on “cultural eutrophication” (1947) helped to guide efforts regarding sewage diversion, fertilizer and manure runoff, and soil erosion from agricultural fields to lakes. He focused much attention on his beloved Lake Mendota, which he could see from his office window at the university. Prior to these efforts much of lake management still revolved around the idea of a lake as a microcosm (Forbes, 1899). John Magnuson, who succeeded Hasler as director of the Laboratory for Limnology, said of Hasler, “He was a big thinker and had grand ideas. He believed you were not done in research until you dealt with its application to society.” During a time when a Washington presence was not in fashion, Hasler spoke out frequently, eloquently and effectively on environmental issues that he knew about and cared about. Hasler was an outstanding scientist, a mentor, a wonderful friend, and an effective spokesman for the protection of natural resources.
Although he had survived four bouts of cancer (colon, lung, skin, and prostate) starting in 1972, all without major chemotherapy, he continued to be active in campus activities until December 2000. He died peacefully in March 2001 at 93.

Er ist’s
[by Eduard Mörike]
Frühling läβt sein blaues Band
Wieder flattern durch die Lüfte;
Süβe, wohlbekannte Düfte
Streifen ahnungsvoll das Land.
Veilchen träumen schon,
Wollen balde kommen.
- Horch, von fern ein leiser Harfenton!
Frühling, ja du bist’s!
Dich hab ich vernommen!

AM PLEASED to acknowledge Hatheway Hasler, Linda Holthaus, John Magnuson, and William Schmitz for details and suggestions; autobiographical materials from the National Academy of Sciences and the University of Wisconsin-Madison; various Limnology News newsletters from the Center for Limnology, University of Wisconsin-Madison; and “Resolution of Respect” in the Bulletin of the Ecological Society of America, July 2001 (S.Carpenter and J.Kitchell; G.E.Likens) in the preparation of this memoir. A transcript of an oral interview by Laura Lord Smail, University of Wisconsin-Madison, Oral History Project, in 1977 was especially helpful.

The publications of Arthur Davis Hasler (John Magnuson)

He wrote and coauthored over 200 peer reviewed scientific papers and authored or contributed to seven books

Hasler, A.D. 1935. The physiology of digestion in plankton Crustacea. I. Some digestive enzymes of Daphnia. Biol. Bull. 68: 207–214.
Hasler, A.D. 1937. Methods for culturing
Daph- nia. pp. 214–215. In: P.S. Galtsoff, F.E. Lutz, P.S. Welch & J.G. Needham (eds), Culture Methods for Invertebrate Animals, Comstock Publication Company, Ithaca, New York.
Hasler, A.D. 1937. The physiology of digestion in plankton Crustacea. II. Further studies on the di- gestive enzymes of (A) Daphnia and Polyphemus; (B) Diaptomus and Calanus. Biol. Bull. 72: 290–298. Galtsoff, P.S., W.A. Chipman Jr., A.D. Hasler & J.B. Engle. 1938. Preliminary report on the cause of the decline of the oyster industry of the York River, VA and the effects of pulp-mill pollution on oysters. US Bureau of Fisheries Investigative Re- port No. 37, U.S. Government Printing Office, Washington. 42 pp.
Hasler, A.D. 1938. Pulp-mill pollution in the York River, VA, and its connection with the decline of the oyster industry. Gamma Alpha Record 28: 44–45. 

Hasler, A.D. 1938. Fish biology and limnology of Crater Lake, Oregon. J. Wildlife Manage. 2: 94–103. Hasler, A.D., R.K. Meyer & H.M. Field. 1939. Spawning induced prematurely in trout with the aid of pituitary glands of the carp. Endocrinology 25: 978–983.
Hasler, A.D., R.K. Meyer & H.M. Field. 1940. The use of hormones for the conservation of muskellunge, Esox masquinongy immaculatus Garrard. Copeia 1: 43–46.
Hasler, A.D. & W.M. Faber. 1941. A tagging method for small fish. Copeia 3: 162–165.
Hasler, A.D. & R.K. Meyer. 1941. Metabolic and reproductive responses of fish to hormones. pp. 403–404.
In: J.G. Needham (ed.) A Symposium on Hydrobiology, University of Wisconsin Press, Madison.

Hasler, A.D. & D.S. Farner. 1942. Fisheries in- vestigations in Crater Lake, Oregon, 1937–1940. J. Wildlife Manage. 6: 319–327.
Hasler, A.D. & R.K. Meyer. 1942. Respiratory responses of normal and castrated goldfish to tel- eost and mammalian hormones. J. Exp. Zool. 91: 391–404.

Nelson, M.N. & A.D. Hasler. 1942. The growth, food, distribution and relative abundance of the fishes of Lake Geneva, Wisconsin in 1941. T. Wisc. Acad. Sci., Arts Lett. 34: 137–148.
Hasler, A.D. & H.F. Deutsch. 1943. Distribution of a vitamin B1 destructive enzyme in fish. P. Soc. Exp. Biol. Med. 53: 63–65.
Andrews, J.D. & A.D. Hasler. 1944. Fluctuations in the animal populations of the littoral zone in Lake Mendota. T. Wisc. Acad. Sci., Arts Lett. 35: 175–186.

Juday, C. & A.D. Hasler. 1944. List of publica- tions dealing with Wisconsin limnology (1871– 1945). T. Wisc. Acad. Sci., Arts Lett. 36: 469–490. Schneberger, E. & A.D. Hasler. 1944. Brule River Survey: Introduction. Wisc. Conservation Bull. 9(8–9): 5–8.
Thomsen, H.P. & A.D. Hasler. 1944. The minnow problem in Wisconsin. Wisc. Conservation Bull. 9(12): 6–8.
Hasler, A.D. 1945. Observations on the winter perch population of Lake Mendota. Ecology 26: 90–94. Hasler, A.D. 1945. Publications of Chancey Juday. Limnol. Soc. Am., Special Publication No. 16: 4–9. Hasler, A.D. 1945. Some recent cooperative re- searches at Wisconsin in fishery biology. Trans.
Tenth North Am. Wildlife Conf. 260–266. Hasler, A.D. 1945. This is the enemy. Science 102: 431.
Hasler, A.D. 1946. A war time view of European biological stations. (An account of some experi- ences and information gathered in Europe, 1945). Biologist 28: 81–93.
Hasler, A.D. 1946. Book review of a ‘Textbook of Limnology’ in German by Franz Ruttner. Ecology 27: 268–269.
Hasler, A.D. & L.V. Whitney. 1946. A combina- tion photo-electric light meter and fish-detector. J. Wildlife Manage. 10: 175–177.
Hasler, A.D., H.P. Thomsen & J.C. Neess. 1946. Facts and comments on raising two common bait minnows. Wisc. Conservation Bull. No. 210-A-46. 14 pp.
Hasler, A.D. 1947. Eutrophication of lakes by domestic drainage. Ecology 28: 383–395.
Galtsoff, P.S., W.A. Chipman Jr., J.B. Engle & H.N. Calderwood. 1947. Ecological and physiolo- gical studies of the effect of sulfate pulp mill wastes on oysters in the York River, VA. Fish. Bull. Fish Wildlife Serve. No. 43: 59–186 (See page 60). Hasler, A.D. 1948. Book review of ‘Duftgelenkte Bienen im Dienste der Landwirtschaft und Imk- erei’ (The Training of Bees for the Benefit of Beekeeping and Agriculture) by K. Von Frisch (ed.). Science 108: 290.
Hasler, A.D. 1948. General Zoology in Labora- tory and Field. William C. Brown, Dubuque, Iowa. Figure 35, p. 53.
Hasler, A.D. & W.G. Einsele. 1948. Fertilization for increasing productivity of natural inland wa- ters. Transactions of the thirteenth North Amer- ican Wildlife Conference, Wildlife Management Institute 527–552.
Hasler, A.D. 1949. Antibiotic aspects of copper treatment of lakes. T. Wisconsin Acad. Sci., Arts Lett. 39: 97–l03.
Hasler, A.D. & J.E. Bardach. 1949. Daily migra- tions of perch in Lake Mendota, Wisconsin. J. Wildlife Manage. 13: 40–51.
Hasler, A.D. & E. Jones. 1949. Demonstrations of the antagonistic action of large aquatic plants on algae and rotifers. Ecology 30: 359–364.
Walker, T.J. & A.D. Hasler. 1949. Detection and discrimination of odors of aquatic plants by the bluntnose minnow (
Hyborhynchus notatus). Phy- siol. Zool. 22: 45–63. 

Hasler, A.D. 1950. Book review of ‘Fortschrittli- che Karpfenteichwirtschaft’ by W. Wunder (ed.). Copeia 2: l59–l60.
Hasler, A.D. & W.J. Wisby. 1950. Use of fish for the olfactory assay of pollutants (phenols) in wa- ter. T. Am. Fish. Soc. 79: 64–70.

Brooks, J.L., G.L. Clarke, A.D. Hasler & L.E. Noland. 1951. Edward Asahel Birge (1851–1950). Arch. Hydrobiol. 45: 235–243.
Hasler, A.D. & W.J. Wisby. 1951. Discrimination of stream odors by fishes and its relation to parent stream behavior. Am. Nat. 85: 223–238.

Hasler, A.D., Brynildson, O.M. & W.T. Helm. 1951. Improving conditions for fish in brown-water bog lakes by alkalization. J. Wildlife Manage. 15: 347–352. Wohlschlag, D.E. & A.D. Hasler. 1951. Some quantitative aspects of algal growth in Lake Mendota. Ecology 32: 581–593.
Brynildson, O.M., A.D. Hasler & J.A. Larsen. 1952. Bog lakes for trout. Wisc. Conservation Bull. 17(11): 1–3.
Hasler, A.D. 1952. Book review of ‘Die Bin- nengewa ̈ sser Bd. XVIII, of Verbreitungsgeschichte der Su ̈ sswassertierwelt Europas’ by A. Thiene- mann. Copeia 2: l20.

Hasler, A.D. 1952. Book review of ‘Die Nahrung der Meerestiere Bd. I, of Handbuch der Seefisc- herei Nordeuropas’ by A. Hagmeier & C. Kunne. Copeia 2: 120.
Nursall, J.R. & A.D. Hasler. 1952. A note on ex- periments designed to test the viability of gametes and the fertilization of eggs by minute quantities of sperm. Prog. Fish Cult. 14: 165–168.
Hasler, A.D. 1953. Chemical sense in animals. In: The Encyclopedia Americana, New York.
Hasler, A.D. 1953. Analysis of factors initiating parent-stream behavior in salmon. pp. 67–81.
In: Proceedings of the Conference on Orientation in Animals, Office of Naval Research, Department of the Navy, Washington D.C.
Hasler, A.D. & J.R. Villemonte. 1953. Observa- tions on the daily movements of fishes. Science 118: 321–322.
Hasler, A.D. & W.J. Wisby. 1953. Salmon, sci- ence, and the sense of smell. Research Reviews, Office Naval Res. June: 11–15.
Hasler, A.D., W.J. Wisby & J.A. Larsen. 1953. Lachse werden umgeleitet. Fischwirt 3: 147–151. Hasler, A.D. 1954. Bog lakes now produce rain- bows. Prog. Fish Cult. 16: 142–143.
Hasler, A.D. 1954. Book review of ‘Die Fisch- dressuren und ihre Sinnesphysiologischen Grund- lagen’ by K. Herter (ed.), Copeia 2: 161.
Hasler, A.D. 1954. Odour perception and orienta- tion in fishes. J. Fish. Res. Board Can. 11: 107–129. Johnson, W.E. & A.D. Hasler. 1954. Rainbow trout production in dystrophic lakes. J. Wildlife Manage. 18: 113–134.
Wisby, W.J. & A.D. Hasler. 1954. Effect of ol- factory occlusion on migrating silver salmon (O. kisutch). J. Fish. Res. Board Can. 11: 472–478. Hasler, A.D. 1955. Participant. In: B. Schaffner (ed.) Group processes: Transactions of the Second Conference, Princeton, N.J. 255 pp.
Hasler, A.D. & J.A. Larsen. 1955. The homing salmon. Sci. Am. 193: 72–75.

Ramaswami, L.S. & A.D. Hasler. 1955. Hormones and secondary sex characteristics in the minnow, Hyborhynchus. Physiol. Zool. 28: 62–68.
Hasler, A.D. 1956. Book review of ‘Die Bin- nengewa ̈ sser in Natur und Kultur.’ Einfu ̈ hrung in die theoretische und angewandte Limnologie (Versta ̈ ndliche Wissenschaft 55) by A. Thiene- mann. Science Monthly 82(4).

Hasler, A.D. 1956. Influence of environmental reference points on learned orientation in fish (Phoxinus). Z. Vergl. Physiol. 38: 303–310. Hasler, A.D. 1956. Isotopes and atomic energy in the service of marine and freshwater agriculture. pp. 103–111. Hearings of 84th Congressional Committee on Atomic Energy. U.S. Government Printing Office, Washington, D.C.
Hasler, A.D. 1956. Laboratory of Hydrobiology, University of Wisconsin. Am. Inst. Biol. Sci. Bull. 5: 31–32.
Hasler, A.D. 1956. Perception of pathways by fishes in migration. Q. Rev. Biol. 31: 200–209. John, K.R. & A.D. Hasler. 1956. Observations on some factors affecting the hatching of eggs and the survival of young shallow-water cisco,
Leucichthys artedi LeSueur in Lake Mendota, Wisconsin. Limnol. Oceanogr. 1: 176–194.

Zicker, E.L., K.C. Berger & A.D. Hasler. 1956. Phosphorus release from bog lake muds. Limnol. Oceanogr. 1: 296–303.
Hasler, A.D. 1957. Natural and artificially (air- ploughing) induced movement of radioactive phosphorus from the muds of lakes. pp. 658–675.
In: UNESCO International Conference on Radioiso- topes, Scientific Research. Vol. IV, Paris, France. Hasler, A.D. 1957. The sense organs: Olfactory and gustatory senses of fishes. pp. 187–209. In: M.E. Brown (ed.) Physiology of Fishes, Vol. II, Academic Press, New York.

Larsen, J.A. & A.D. Hasler. 1957. Heimkehrer ‘Lachs.’ Fischerei-Zeitung 19: 370–372.
Hasler, A.D. 1958. Book review of a ‘Treatise on Limnology’ by G.E. Hutchinson. Science 127: 88. Hasler, A.D. 1958. Perception of pathways by fish in migration. pp. 451–467.
In: A.A. Buzzati- Traverso (ed.) Perspectives in Marine Biology, University of California Press, Berkeley.

Hasler, A.D. 1958. Sun and odor orientation in migrating fishes. Abstracts of the XVth Interna- tional Congress of Zoology, Section X, paper 19. Hasler, A.D. & W.J. Wisby. 1958. Perch and lake
research on Mendota. Wisc. Conservation Bull. 23(3): 1–5.
Hasler, A.D. & W.J. Wisby. 1958. The return of displaced largemouth bass and green sunfish to a ‘home’ area. Ecology 39: 289–293.

Hasler, A.D., R.M. Horrall, W.J. Wisby & W. Braemer. 1958. Sun-orientation and homing in fishes. Limnol. Oceanogr. 3: 353–361.
Schmitz, W.R. & A.D. Hasler. 1958. Artificially induced circulation of the lake by means of com- pressed air. Science 128: 1088–1089.

Hasler, A.D. 1959. Method of repelling fish com- prising treating with potassium phenyl acetate. U. S. Patent Office, #2880133: 6 pp.
Parker, R.A. & A.D. Hasler. 1959. Movements of some displaced centrarchids. Copeia 1: 11–18. Hasler, A.D. 1960. Guideposts of migrating fishes. Science 132: 785–792.

Hasler, A.D. 1960. Homing orientation in migrating fishes. Erg. Biol. 23: 94–115.
Hasler, A.D. & H.O. Schwassmann. 1960. Sun orientation in fish at different latitudes. Cold Spring Harb. Symp. 25: 429–441.

Likens, G.E. & A.D. Hasler. 1960. Movement of radiosodium in a chemically stratified lake. Science 131: 1676–1677.
Schneider, H. & A.D. Hasler. 1960. Laute und Lauterzeugung beim Susswassertrommler
Aplodi- notus grunniens Rafinesque (Sciaenidae, Pisces). Z. Vergl. Physiol. 43: 499–517.

Stross, R.G. & A.D. Hasler. 1960. Some lime-in- duced changes in lake metabolism. Limnol. Oceanogr. 5: 265–272.
Hasler, A.D. 1961. Arboretum serves in study of fish migration. Arboretum News 10(1).

Hasler, A.D. 1961. Feier zur Einweihung des Neubaues der hydrogiologischen Anstalt der Max- Planck Gesellschaft. MittTeilungen aus der Planck- Geschellschaft zur forderung des Wis- senschaften 5: 314–316.
Hasler, A.D. 1961. The organism and the envi- ronment. pp. 65–77. In: Marguerite Gilstrap (ed.) Promise of the Life Sciences, U.S. Department of Agriculture Graduate School.
McNaught, D.C. & A.D. Hasler. 1961. Surface schooling and feeding behavior in the white bass, Roccus chrysops (Rafinesque), in Lake Mendota. Limnol. Oceanogr. 6: 53–60.
Stross, R.G., J.C. Neess & A.D. Hasler. 1961. Turnover time and production of the planktonic
crustacea in limed and reference portion of a bog lake. Ecology 42: 237–245.
Hasler, A.D. 1962. Limnologists and oceanogra- phers discuss lakes, rivers, and aquatic life. Science 138: 698–703.

Hasler, A.D. 1962. Universita ̈ t und Forschung: Die Universita ̈ t von Wisconsin. Naturwissens- chaftliche Rundschau 14: 150–152.
Hasler, A.D. 1962. Wegweiser fu ̈ r Zugfische. Na- turwissenschaftliche Rundschau 15: 302–310. Likens, G.E. & A.D. Hasler. 1962. Movements of radiosodium (Na
24) within an ice-covered lake. Limnol. Oceanogr. 7: 48–56.

Hasler, A.D. 1963. Discussant on ‘Components of ecosystems, Phytoplankton Productivity, and Trophic change: experimental approaches.’ In: G.A. Riley (ed.) Proceedings of the first interdis- ciplinary conference on marine biology, American Institute of Biological Sciences, Washington D.C. Hasler, A.D. 1963. Experimentelle Limnologie in der heutigen Wasserforschung. Wasser und Ab- wasser 1–2: 19–23.
Hasler, A.D. 1963. The Laboratory of Limnology and associated field units of the University of Wisconsin. Am. Zool. 3: 337–340.
Hasler, A.D. 1963. The Laboratory of Limnology at the University of Wisconsin. Wisc. Acad. Rev. 10(1): 1–7.

Hasler, A.D. 1963. Wisconsin 1940–1961. Chapter 2, pp. 55–93. In: David Frey (ed.) Limnology in North America, University of Wisconsin Press, Madison.
Hasler, A.D. & H.F. Henderson. 1963. In- strumentation problems in the study of homing in fish. Bio-Telemetry Conference pp. 195–201. Hasler, A.D. & G.E. Likens. 1963. Biological and physical transport of radio-nuclides in stratified lakes. pp. 463–470. In: V. Schultz & A. Klement (eds), Radioecology, Reinhold Publishers, New York. Hasler, A.D. 1964. Experimental limnology. BioScience 14: 36–38.
McNaught, D.C. & A.D. Hasler. 1964. Rate of movement of population of Daphnia in relation to changes in light intensity. J. Fish. Res. Board Can. 21: 291–318.
Schwassmann, H.O. & A.D. Hasler. 1964. The role of the sun’s altitude in sun orientation of fish. Physiol. Zool. 37: 163–178.
Gammon, J.R. & A.D. Hasler. 1965. Predation by introduced muskellunge on perch and bass, I:
Years 1–5. T. Wisc. Acad. Sci., Arts and Lett. 54: 249–272.
Hasler, A.D., H.F. Henderson, R.M. Horrall & E.S. Gardella. 1965. Orientation of homing white bass. Am. Zool. 5: 703.

Hunter, J.E. & A.D. Hasler. 1965. Spawning asso- ciation of the redfin shiner, Notropis umbratilis, and green sunfish, Lepomis cyanellus. Copeia 3: 265–281. Rabinowitch, V. & A.D. Hasler. 1965. The Inter- national Biological Program. Bull. Atom. Sci. 21: 32–34.
Hasler, A.D. 1966. Underwater Guideposts – Homing of Salmon. University of Wisconsin Press, Madison. 155 pp.
Henderson, H.F., A.D. Hasler & G.G. Chipman. 1966. An ultrasonic transmitter for use in studies of movements of fishes. T. Am. Fish. Soc. 95: 350– 356.

Hergenrader, G.L. & A.D. Hasler. 1966. Diel ac- tivity and vertical distribution of yellow perch (Perca flavescens) under the ice. J. Fish. Res. Board Can. 23: 499–509.
Malueg, K.W. & A.D. Hasler. 1966. Echo sounder studies on diel vertical movements of larvae in Wisconsin (U.S.A.) lakes. Verhandlungen Inter- national Verein Limnologie 16: 1697–1708. McNaught, D.C. & A.D. Hasler. 1966. Photo- environments of planktonic Crustacea in Lake Michigan. Verhandlungen International Verein Limnologie 16: 194–203.
Schwartz, E. & A.D. Hasler. 1966. Perception of surface waves by the black-stripe topminnow, Fundulus notatus. J. Fish. Res. Board Can. 23: 1331–1352.
Schwartz, E. & A.D. Hasler. 1966. Superficial lateral line sense organs of the mudminnow (Umbra limi). Z. Vergl. Physiol. 53: 317–327. Corey, R.B., A.D. Hasler, G.F. Lee, F.H. Schraufnagel & T.L. Wirth. 1967. Excessive water fertilization. Report to Water Subcommittee Nat- ural Resources Committee, State Agencies, Wis- consin. 50 pp.
Hasler, A.D. 1967. Can ecology provide the basis for synthesis? Discussion on: pp. 41–45. In: M.E. Garney & J.R. Hibbs (eds), Social Sciences and the Environment, University of Colorado Press, Boulder.
Hasler, A.D. 1967. Das Ratsel der Lachswander- ungen. Naturwissenschaft und Medizin 4(17): 20–31. Hasler, A.D. 1967. Underwater guideposts for
migrating fishes. pp. 1–20. In: R.M. Storm (ed.) Animal Orientation and Navigation, Proceedings of the Twenty-Seventh Annual Biology Collo- quium, Oregon State University Press, Corvallis. Hasler, A.D. 1967. Technology and man’s relation to his natural environment. pp. 158–167. In: C.P. Hall (ed.) Human Values and Advancing Tech- nology, Friendship Press, New York.
Hasler, A.D. & M.E. Swenson. 1967. Eu- trophication. Science 158: 278–282.
Hasler, A.D. & M.E. Swenson. 1967. Home from the sea. New Sci. 12: 96–98.

Hergenrader, G.L. & A.D. Hasler. 1967. Seasonal changes in swimming rates of yellow perch in Lake Mendota as measured by sonar. T. Am. Fish. Soc. 96: 373–382.
Wright, T.D. & A.D. Hasler. 1967. An electro- phoretic analysis of the effects of isolation and homing behavior upon the serum proteins of the white bass (Roccus chrysops) in Wisconsin. Am. Nat. 101: 401–413.
Hasler, A.D. 1968. Memory in homing of migra- tory fishes. pp 247–255. In: D. Ingle (ed.) The Central Nervous System and Fish Behavior, Uni- versity of Chicago.
Hasler, A.D. & B. Ingersoll. 1968. Dwindling lakes. Nat. Hist. 77: 8–31.
Hergenrader, G.L. & A.D. Hasler. 1968. Influence of changing seasons on schooling behavior of yellow perch. J. Fish. Res. Board Can. 25: 711– 716.

Wissing, T.E. & A.D. Hasler. 1968. Calorific va- lues of some invertebrates in Lake Mendota, Wisconsin. J. Fish. Res. Board Can. 25: 2515– 2518.
Hasler, A.D. 1969. Acting responsibly to our environment, a profoundly human issue. INT- ECOL Bull. 1: 2–12.
Hasler, A.D. 1969. Book review of the ‘Sockeye Salmon (
Oncorhynchus nerka)’ by R.E. Foerster. Q. Rev. Biol. 44: 324–325.

Hasler, A.D. 1969. Cultural eutrophication is reversible. BioScience 19: 425–431.
Hasler, A.D., E.S. Gardella, R.M. Horrall & H.F. Henderson. 1969. Open water orientation of white bass,
Roccus chrysops, as determined by ultrasonic tracking methods. J. Fish. Res. Board Can. 26: 2173–2192.

Hasler, A.D., R.M. Horrall, A.B. Stasko & E.S. Gardella. 1969. Following migrating fishes equip-
ped with ultrasonic transmitters. Verhandlungen International Verein Limnologie 19: 516.
Parker, M. & A.D. Hasler. 1969. Studies on the distribution of cobalt in lakes. Limnol. Oceanogr. 14: 229–241.

Sager, P. & A.D. Hasler. 1969. Species diversity in lacustrine phytoplankton. I. The components of the index of diversity from Shannon’s formula. Am. Nat. 103: 51–59.
Hasler, A.D. 1970. Chemical ecology of fish. pp. 219–234. In: E. Sondheimer & J.B. Simeone (eds), Chemical Ecology, Academic Press, New York. Hasler, A.D. 1970. Man-induced eutrophication of lakes. pp. 110–125. In: S.F. Singer (ed.) Global Effects of Environmental Pollution.
Hasler, A.D. 1970. What is ecology? pp. 431–473. In: Philip Handler (ed.) Biology and the Future of Man, Oxford University Press, New York. Hasler, A.D. & J.J. Tibbles. 1970. A study of depth distribution of perch (Perca flavescens) with a rolling gill net. Betrage Deutsche Wis- senschaften Kommunicationen Meeresforschung 21: 46–55.
Hasler, A.D., R.M. Horrall, A.B. Stasko & A.E. Dizon. 1970. Orientation cues and tracking of salmonid fishes. Proc. Natl. Acad. Sci. 66: 13–14. Hasler, A.D. 1971. Eutrophication, cultural. McGraw Hill Encyclopedia of Science and Tech- nology, New York, pp.136–138.
Hasler, A.D. 1971. Man in the Living Environ- ment. The Institute of Ecology Report of the Workshop on Global Ecological Problems, Uni- versity of Wisconsin Press, Madison. 267 pp. Hasler, A.D. 1971. Orientation and fish migration. pp. 429–510. In: W.S. Hoar & D.J. Randall (eds), Physiology of Fishes, Vol. VI, Academic Press, New York.
Larsen, J.A. & A.D. Hasler. 1971. Giving our lakes a chance. pp. 138–145. In: Science Year, The World Book Science Annual 1971, Field Enter- prises Educational Corporation.
Madison, D.M. & A.D. Hasler. 1971. Recent ad- vances in salmon orientation. Am. Zool. 11: 629. Madison, D.M. & A.D. Hasler. 1971. Modified float and hybrid tracking systems. Underwater Telemetry Newsletter 1(2): 14–17.
Wissing, T.E. & A.D. Hasler. 1971. Effects of swimming activity and food intake on the re- spiration of young-of-the-year white bass, Morone chrysops. T. Amer. Fish. Soc. 100: 537–543.
Wissing, T.E. & A.D. Hasler. 1971. Intraseasonal change in caloric content of some freshwater in- vertebrates. Ecology 52: 371–373.
Chidambaram, S., R.K. Meyer & A.D. Hasler. 1972. Effects of hypophysectomy, pituitary auto- grafts, prolactin, temperature and salinity of the medium on survival and natremia in the bullhead,
Ictalurus melas. Comp. Biochem. Physiol. 43A: 443–457.

Cooper, J.C. & A.D. Hasler. 1972. Electro- encephalography (EEG) and salmon homing: a continuing controversy. Am. Zool. 12: 653. Hasler, A.D. 1972. Review of analysis of ecosys- tems of US-IBP. pp. 893–895. In: Z. Kajak & A. Hillbricht-Ilkowska (eds), Proceedings of the IBP- UNESCO Symposium on Productivity Problems of Freshwaters, Kazimierz Dolny, Poland. Hasler, A.D. 1972. A zoologist’s responsibility to our living environment. Am. Zool. 12: 9–11. Kaya, C.M. & A.D. Hasler. 1972. Photoperiod and temperature effects on the gonads of green sunfish, Lepomis cyanellus (Rafinesque), during the quiescent, winter phase of its annual sexual cycle. T. Am. Fish. Soc. 101: 270–275.
Madison, D.M., A.D. Hasler & G.G. Chipman. 1972. A miniature, 5-channel ultrasonic transmit- ter. Underwater Telemetry 2: 3.
Madison, D.M., A. Scholz & A.D. Hasler. 1972. Behavioral evidence of ‘imprinting’ to chemical cues in salmon (motion picture). Am. Zool. 12: 643–644.

Madison, D.M., R.M. Horrall, A.B. Stasko & A.D. Hasler. 1972. Migratory movements of adult sockeye salmon (Oncorhynchus nerka) in coastal British Columbia as revealed by ultrasonic track- ing. J. Fish. Res. Board Can. 29: 1025–1033. Scholz, A., D.M. Madison, A.B. Stasko, R.M. Horrall & A.D. Hasler. 1972. Orientation of sal- mon in response to currents in or near the home stream. Am. Zool. 12: 654.
Stewart, K.M. & A.D. Hasler. 1972. Limnology of some Madison lakes: annual cycles. T. Wisc. Acad. Sci., Arts Lett. 60: 87–123.
Teraguchi, M., T. Wissing & A.D. Hasler. 1972. Change in caloric content of adult males of
Mysis relicta (Loven) during a diel migratory cycle. Am. Mid. Nat. 88: 235–239.

Baumann, P.C., A.D. Hasler, J.F. Koonce & M. Teraguchi. 1973. Biological investigations of Lake Wingra. U.S. Environmental Protection Agency,
Washington, D.C. Ecology Research Series EPA- R3-73-044. 119 pp.
Chidambaram, S., R.K. Meyer & A.D. Hasler. 1973. Effects of hypophysectomy, isletectomy and ACTH on glycemia and hematocrit in the bull- head,
Ictalurus melas. J. Exp. Zool. 184: 75–80. Cooper, J.C. & A.D. Hasler. 1973. An electro- physiological approach to salmon homing. Fish- eries Research Board of Canada Technical Report No. 415. Pacific Biological Station, Nanaimo, British Columbia. 44 pp.

Dizon, A., R. Horrall & A.D. Hasler. 1973. Long- term olfactory ‘memory’ in coho salmon, On- corhynchus kisutch. Fish. Bull. 71: 315–317. Dizon, A.E., R.M. Horrall & A.D. Hasler. 1973. Olfactory electroencephalographic responses of homing coho salmon, Oncorhynchus kisutch, to water conditioned by conspecifics. Fish. Bull. 71: 893–896.
Hasler, A.D. 1973. Causes and correctives of man- made eutrophication. pp. 141–163. In: H. Sioli (ed.) Sonderdruck aus o ̈ kologie und lebensshutz in internationaler sicht, Verlag Rombach and Com- pany, Freiburg, Germany.
Hasler, A.D. 1973. Foreword. pp. ix-x. In: R.M. Darnell (ed.) Ecology and Man, Wm. C. Brown Publishers, Iowa.
Hasler, A.D. 1973. Poisons, phospates, preserva- tion, people and politics – a fish eye’s view of ecology. T. Am. Fish. Soc. 102: 213–224. Madison, D.M., A.T. Scholz, J.C. Cooper, R.M. Horrall, A.D. Hasler & A.E. Dizon. 1973. I. Ol- factory hypotheses and salmon migration: a sy- nopsis of recent findings. Fisheries Research Board of Canada Technical Report No. 414. Pacific Bio- logical Station, Nanaimo, British Columbia. 35 pp. Scholz, A.T., J.C. Cooper, D.M. Madison, R.M. Horrall, A.E. Dizon & R.J. Poff. 1973. Olfactory imprinting in coho salmon: behavioral and elec- trophysiological evidence. Proceedings of the 16th Conference of Great Lakes Research: 143–153. Stasko, A.B., R.M. Horrall, A.D. Hasler & D. Stasko. 1973. Coastal movements of mature Fra- ser River pink salmon (
Oncorhynchus gorbuscha) as revealed by ultrasonic tracking. J. Fish. Res. Board Can. 30: 1309–1316.

Cooper, J.C. & A.D. Hasler. 1974. Electro- encephalographic evidence for retention of olfac- tory cues in homing coho salmon. Science 183: 336–338.
Hasler, A.D. 1974. The Institute of ecology. Bull. Ecol. Soc. Am. 55(2): 7–9.
Hasler, A.D. 1974. Review of ‘Salmon: Their Fight for Survival’ by A. Netboy. T. Am. Fish. Soc. 103: 835–836.

Hasler, A.D. 1974. Unification of land-water eco- logical systems. Presidential Address. pp. 4–7. In: Structure, Functioning and Management of Eco- systems: Proceedings of the First International Congress of Ecology, The Hague, The Netherlands. Hasler, A.D. & J.A. Larson. 1974. The homing salmon. pp. 52–55. In: D.R. Griffin (compiler) Animal Engineering: Readings from Scientific American, W.H. Freeman and Company, San Francisco.
Cooper, J.C. & A.D. Hasler. 1975. Morpholine as olfactory stimulus in fish. Science 187: 81–82. Cooper, J.C. & A.D. Hasler. 1975. Olfactory im- printing and memory in salmonids. Am. Zool. 15: 811. Gallepp, G.W. & A.D. Hasler. 1975. Behavior of larval caddisflies (Brachycentrus spp.) as influenced by marking. Am. Mid. Nat. 93: 247–254.
Hasler, A.D. (ed.). 1975. Coupling of Land and Water Systems, Springer-Verlag, New York. 309 pp.
Hasler, A.D. 1975. How the salmon comes home. pp. 184–191.
In: Science Year, The World Book Science Annual, Field Enterprises Educational Corporation, Chicago.

Hasler, A.D. 1975. Man-induced eutrophication of lakes. pp. 383–399. In: S.F. Singer (ed.) The Changing Global Environment, D. Reidel Pub- lishers, Dordrecht-Holland.
Hasler, A.D. & R. Lanier. 1975. The International Association for Ecology – A brief account of the development process and of current activities. INTECOL Bull. 5: 3–17.
Scholz, A.T., R.M. Horrall, J.C. Cooper, A.D. Hasler, D.M. Madison, R.J. Poff & R.I. Daly. 1975. Artificial imprinting of salmon and trout in Lake Michigan. Wisconsin Department of Natural Resources and Fish Management Report 80. 46 pp. Teraguchi, M., A.D. Hasler & A.M. Beeton. 1975. Seasonal changes in the response of Mysis relicta Loven to illumination. Verhandlungen Interna- tional Verein fu ̈ r Limnologie 19: 2989–3000. Cooper, J.C. & A.D. Hasler. 1976. Electrophysio- logical studies of morpholine-imprinted coho salmon (Oncorhynchus kisutch) and rainbow trout (Salmo gairdneri). J. Fish. Res. Board Can. 33: 688–694.
Cooper, J.C., A.T. Scholz, R.M. Horrall, A.D. Hasler & D.M. Madison. 1976. Experimental con- firmation of the olfactory hypothesis with homing, artificially imprinted coho salmon (Oncorhynchus kisutch). J. Fish. Res. Board Can. 33: 703–710. Gasith, A. & A.D. Hasler. 1976. Airborne litterfall as a source of organic matter in lakes. Limnol. Oceanogr. 21: 253–258.
Hasler, A.D. 1976. Review of ‘Marine Ecology: A comprehensive integrated treatise on life in oceans and coastal waters; Vol. II: Physiological mecha- nisms – Part 2: Orientation‘ by Otto Kinne (ed.). T. Am. Fish. Soc. 105: 183–185.
Hasler, A.D. & J.C. Cooper. 1976. Chemical cues for homing salmon. Experientia 32: 1091–1093. Scholz, A.T., R.M. Horrall, J.C. Cooper & A.D. Hasler. 1976. Imprinting to chemical cues: the basis for home stream selection in salmon. Science 192: 1247–1249.
Stasko, A.B., R.M. Horrall & A.D. Hasler. 1976. Coastal movements of adult Fraser River sockeye salmon (Oncorhynchus nerka) observed by ultrasonic tracking. T. Am. Fish. Soc. 105: 64–71.
Johnsen, P.B. & A.D. Hasler. 1977. Winter ag- gregations of carp (Cyprinus carpio) as revealed by ultrasonic tracking. T. Am. Fish. Soc. 106: 556–559. Hasler, A.D. & A.T. Scholz. 1978. Olfactory im- printing in coho salmon. pp 356–369. In: K. Schmidt- Koenig & W.T. Keeton (eds), Animal Migration, Navigation, and Homing, Springer-Verlag, Berlin. Hasler, A.D., A.T. Scholz & R.M. Horrall. 1978. Olfactory imprinting and homing in salmon. Am. Sci. 66: 347–355.
Scholz, A.T., J.C. Cooper, R.M. Horrall & A.D. Hasler. 1978. Homing of morpholine-imprinted brown trout, Salmo trutta. Fish. Bull. 76: 293–295. Scholz, A.T., C.K. Gosse, J.C. Cooper, R.M. Horrall, A.D. Hasler, R.I. Daly & R.J. Poff. 1978. Homing of rainbow trout transplanted in Lake Michigan: a comparison of three procedures used for imprinting and stocking. T. Am. Fish. Soc. 107: 439–443.
Hasler, A.D. 1979. Enlightened management of aquatic biota. pp. 109–121. In: N. Polunin (ed.) Growth Without Ecodisasters, Macmillan Press, England.
Hasler, A.D. 1979. Book review of ‘Pacific Salmon and Steelhead Trout’ by R.J. Childerhose & Marj Trim. Fish., Bull. Am. Fish. Soc. 5(2): 44–45.
Hasler, A.D. & A.T. Scholz. 1980. Artificial im- printing: a procedure for conserving salmon stocks. pp. 179–199. In: J.E. Bardach, J.J. Mag- nuson, R.C. May & J.M. Reinhart (eds), Fish behavior and its use in the capture and culture of fishes, Physiological and behavioral manipulation of food fish as production and management tools; 3–8 November, 1977; Bellagio, Italy. International Center for Living Aquatic Resources Manage- ment, Manila, Philippines.
Johnsen, P.B. & A.D. Hasler. 1980. The use of chemical cues in the upstream migration of coho salmon, Oncorhynchus kisutch Walbaum. J. Fish Biol. 17: 67–73.
Hasler, A.D. 1982. Book review of ‘Fishes: An Introduction to Ichthyology’ by P.B. Moyle & J.J. Cech, Jr. BioScience 32: 753.
Hasler, A.D. 1982. Causas y correccion de la eu- trofizacion debida al hombre. pp 139–163.
In: H. Sioli (ed.) Ecologia y Proteccion de la Naturaleza, Conclusiones Internacionales, Editorial Blume, Barcelona, Spain.

Hasler, A.D. 1982. Preface: ‘Fische und ihr Ver- halten.’ pp. 1–4. In: Die Erforschung der G‘eheimnisvollen Welt unter Wasser. G.K.H. Zu- panc (ed.) Tetra Verlag.
Hasler, A.D. 1983. Art Hasler in China. Hom- opiscis rusticus: Newsletter for retired members of the American Fisheries Society, Bethesda, Mary- land 2(3–4): 7–10.
Hasler, A.D. 1983. Synthetic chemicals and pher- omones in homing salmon. pp. 103–116. In: J.C. Rankin, T.J. Pitcher & R. Duggan (eds), Control Processes in Fish Physiology, Croom Helm, London.
Hasler, A.D. & A.T. Scholz. 1983. Olfactory Imprinting and Homing in Salmon. Springer- Verlag, Berlin, 134 pp.
Hasler, A.D. 1984. Editorial – How lucky can you be? J. Great Lakes Res. 10: 333.

asler, A.D. 1985. Preface to ‘Fish and Their Behavior’ by G.K.H. Zupanc. Tetra Press, Germany. 188 pp. Hasler, A.D. 1985. Book review of ‘Fish Migra- tion’ by B.A. McKeown. Croom Helm, London. Zeitschrift fu ̈ r Tierpsychologie 69: 168–169.
Hasler, A.D. 1985. Book review of ‘The Control of Fish Migration’ by R.J.F. Smith, Springer-Verlag, Berlin. Zeitschrift fu ̈ r Tierpsychologie 70: 11–12. Hasler, A.D. 1996. Potential application of imprinting to the rehabilitation of salmon stocks in rivers with international borders. pp. 83–86. In: R.A. Neal (ed.) International Development, Proceedings of the World Fisheries Congress, Theme 4, New Delhi, India.
Beckel, A.L. 1987. Breaking New Waters, A Century of Lim- nology at the University of Wisconsin. T. Wisc. Acad. Sci., Arts Lett., Special Issue. 122 pp.
Carpenter, S. & J. Kitchell. 2001. Resolution of respect, Arthur Davis Hasler, 1908–2001. Bull. Ecol. Soc. Am. 82: 172–173.
Hasler, A.D., R.M. Horrall, W.J. Wisby & W. Braemer. 1958. Sun-orientation and homing in fishes. Limnol. Oceanogr. 3: 353–361.
Hasler, A.D. & W.J. Wisby. 1951. Discrimination of stream odors by fishes and its relation to parent stream behavior. Am. Nat. 85: 223–238.
Likens, G.E. 2002. Arthur Davis Hasler 1908–2001. Bio- graphical Memoirs, vol. 82. National Academy Press, Washington D.C. pp. 1–14.
Madison, D.M., R.M. Horrall, A.B. Stasko & A.D. Hasler. 1972. Migratory movements of adult sockeye salmon (Oncorhynchus nerka) in coastal British Columbia as revealed by ultrasonic tracking. J. Fish. Res. Board Can. 29: 1025–1033.
Magnuson, J.J. 2002. Three generations of limnology at the University of Wisconsin-Madison. Verhandlungen Interna- tional Verein fu ̈ r Limnologie 28: 856–860.
Mattmiller, B. & J. Magnuson. 2001. Arthur Hasler, limnology pioneer. Fisheries 26: 32.
Mattmiller, B. & J.F. Kitchell. 2002. Obituary, Arthur Hasler, 1908–2001. J. Fish Biol. 60: 1–2.
Scholz, A.T., R.M. Horrall, J.C. Cooper & A.D. Hasler. 1976. Imprinting to chemical cues: the basis for home stream selection in salmon. Science 192: 1247–1249.
Schwassmann, H.O. & W. Braemer. 1961. The effect of exper- imentally changed photoperiod on the sun-orientation rhythm of fish. Physiol. Zool. 34: 273–286.
Wisby, W.J. & A.D. Hasler. 1954. Effect of olfactory occlusion on migrating silver salmon (O. kisutch). J. Fish. Res. Board Can. 11: 472–478. 


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  3. Hi, I am Erwin Roth, LDS Historian Salzburg Austria. Collect history of LDS 1900 - 1955 in Salzburg. Great Blog.

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    Would love to learn more about Arthur D.Hasler as missionary and 1945 as Expert in relation to the branch of Salzburg. Best greetings Erwin

  4. As a Hasler myself, I'm honoured to be somehow related to such a man!
    Many thanks.
    Scott Hasler, London, UK

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  7. Hello Dr. Hasler,

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